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Matrix 13269 of Study 12851

About Citation title: "Phylogenetic relationships of the suckermouth armoured catfishes (Loricariidae) with emphasis on the Hypostominae and the Ancistrinae".
About Study name: "Phylogenetic relationships of the suckermouth armoured catfishes (Loricariidae) with emphasis on the Hypostominae and the Ancistrinae".
About This study is part of submission 12851 (Status: Published).

Matrices

Title: Loricariidae

Rows

Taxon Label Row Segments Characters 1?–30
Acanthicus hystrix  (none) 100000220110010111010010001001
Ancistrus pirareta  (none) 001000220100010111011000021011
Ancistrus sp.  (none) 001000220100010111011000021011
Astroblepus spp. 4  (none) 100000000000000000011011000000
Baryancistrus niveatus  (none) 101000220100010111010000021001
Chaetostoma anomala  (none) 010001220100011111011000020001
Chaetostoma pearsei  (none) 010001220100011111011000020001
Chaetostoma platyrhynchus  (none) 010001220110011111011000020001
Chaetostoma sovichthys  (none) 010001220100011111011000020001
Chaetostoma stannii  (none) 010001220100011111011000020001
Cordylancistrus torbesensis  (none) 000001220000010111000000020011
Corydoras spp. 4  (none) 000000000000000000000000011000
Corymbophanes andersoni  (none) 101000210000011111000000011010
Corymbophanes kaiei  (none) 101000210000011111000000011000
Crossoloricaria sp.  (none) 000100001111120000000000010100
Crossoloricaria venezuelae  (none) 000100001111120000000100010100
Dekeyseria pulcher  (none) 001000220100010111010000001001
Dekeyseria scaphirhyncha  (none) 0010002201{01}0010111000000001
Delturus anguillicaudatus  (none) 101000110010010001010010000111
Dianema longibarbis  (none) 100000000000000000000000011000
Dolichancistrus cobrensis  (none) 000001210100010111010000020011
Dolichancistrus pediculatus  (none) 0100012201000101110110000200{0
Exastilithoxus fimbriatus  (none) 0110?0110000000001000010001000
Exastilithoxus sp.  (none) 0?1000110000000001011010001000
Glyptoperichthys gibbiceps  (none) 1010002101{01}0010011000000002
Glyptoperichthys lituratus  (none) 101000210100010011000000002001
Glyptoperichthys punctatus  (none) 001000220100010011010000002001
Harttia sp.  (none) 100000110110000011010000010111
Hemiancistrus holostictus  (none) 101000220110010011010000021001
Hemiancistrus landoni  (none) 101000220110010011010000021011
Hemiancistrus maracaiboensis  (none) 101000220110010011010000021001
Hemiancistrus megacephalus  (none) 001000220110000111011000021001
Hemiancistrus panamensis  (none) 101000220110010011010000021001
Hemiancistrus sp.  (none) 000000220100010111010000021001
Hemiancistrus sp. Brazil  (none) ??????????????????????????????
Hemipsilichthys  (none) 102100210000000111010010002011
Hemipsilichthys bahianus  (none) 102000210000010111010010001011
Hemipsilichthys cameroni  (none) 101000210000010111010000001011
Hemipsilichthys nudulus  (none) 102000110000010111010011001011
Hemipsilichthys sp.  (none) 102000210000010111011000001011
Hemipsilichthys splendens  (none) 102000210000010111010000001011
Hisonotus sp.  (none) ??????????????????????????????
Hoplosternum littorale  (none) 100000000000000010000000011000
Hypancistrus inspector  (none) ??????????????????????????????
Hypancistrus zebra  (none) 101000220110010111010000021011
Hypoptopoma sp.  (none) 002000110100001111000000002011
Hypostomus albopunctatus  (none) 101000220100010011010000021001
Hypostomus ammophilus  (none) 002100210100010111000011002001
Hypostomus bolivianus  (none) 101000220110010011010000021001
Hypostomus boulengeri  (none) 101000220100010011010000121001
Hypostomus cochliodon  (none) 101000220100011011010000{01}21
Hypostomus commersoni  (none) 002000220100010011000000121001
Hypostomus cordovae  (none) 101000220100010011010000121001
Hypostomus emarginatus 1  (none) 001000220100011111000010021001
Hypostomus emarginatus 2  (none) 001000220100011111000010021001
Hypostomus francisci  (none) 101000220100010011010000{01}21
Hypostomus hemicochliodon  (none) 101000220110010011010000121001
Hypostomus hondae  (none) 100000220100010011010000121001
Hypostomus micromaculatus  (none) 101000220110010011010000121001
Hypostomus plecostomoides  (none) 101000220100010011010000121001
Hypostomus plecostomus  (none) 101000220100010011010000121001
Hypostomus plecostomus 2  (none) 101000220100010011010000121001
Hypostomus punctatus  (none) 101000220110010011010000021001
Hypostomus robinii  (none) 100000220110010011010000121001
Hypostomus sp. Round Snout 1  (none) 101000220110010011010000021001
Hypostomus sp. Round Snout 2  (none) 101000220100010011010000121001
Hypostomus spinosissimus  (none) ??????????????????????????????
Hypostomus squalinus  (none) 001000220100011111000010012001
Hypostomus unicolor  (none) 002100210100010111000011002001
Isbrueckerichthys alipionis  (none) 002100110000010111000000001011
Isbrueckerichthys duseni  (none) 002100210000010111000000001011
Ixinandria montebelloi  (none) 000000000000000001000010010100
Kronichthys sp. 1  (none) 102000110000010111010000002011
Kronichthys sp. 2  (none) 102000210000010111010010002011
Lamontichthys llanero  (none) 010000110110000011000000010101
Lasiancistrus maracaiboensis  (none) 001010220110010111011000021011
Lasiancistrus sp.  (none) 001010220110010111011000021011
Leporacanthicus galaxias  (none) 100000110000010101010010001000
Leptoancistrus canensis  (none) 110001220000011111011000020001
Liposarcus multiradiatus  (none) 002000210100010011010000{01}02
Liposarcus pardalis  (none) 002000210100010011010000002001
Lithogenes villosus  (none) 00??00000000000000000000001100
Lithoxancistrus orinoco  (none) 0100102201{01}0010111011000020
Lithoxus bovallii  (none) 002000110000000001000010001000
Lithoxus lithoides  (none) 002000110000000001000010001000
Loricaria cataphracta  (none) 000000001111120000000100010100
Loricariichthys brunneus  (none) 000000001100020001000100110100
Megalancistrus gigas  (none) 100000210110010011010000011001
Microlepidogaster sp.  (none) 002000110000001111000000002011
Nannoptopoma spectabile  (none) 002000110000001111000010002000
Neblinichthys pilosus  (none) 101000220100011111010000021011
Neblinichthys roraima  (none) 10100022010001?1110100000???11
Neoplecostomus microps  (none) 002000110000000111000010001000
Neoplecostomus paranensis  (none) 002000110000000111000010001000
Otocinclus vestitus  (none) 002000110110011111000000002011
Otocinclus vittatus  (none) 002000210110011111000000002011
Panaque albomaculatus  (none) 001000220110011111010000020011
Panaque maccus  (none) 001000220110011111010000021011
Panaque nigrolineatus  (none) 000000220100011111000000021001
Panaque pariolispos  (none) 100000220110011011010000021001
Parancistrus aurantiacus  (none) 100000220110010011010100011011
Pareiorhina sp.  (none) 102000110000010111010010002011
Pareiorhina rudolphi  (none) 102000110000000111010010001011
Parotocinclus britskii  (none) 002000110000001111000000002011
Peckoltia oligospila  (none) 102000220110010111010000021011
Peckoltia sp. 1  (none) 100000220110011111010000021011
Peckoltia sp. 2  (none) 100000220110011111010000021011
Peckoltia sp. Big Spot  (none) ??????????????????????????????
Peckoltia ucayalensis  (none) 001000220110011011000000021011
Pogonopoma parahybae  (none) 002000220100010011010000101001
Pogonopoma wertheimeri  (none) 102000220100010011110000101001
Pseudacanthicus leopardus  (none) 100000210100011011010000011001
Pseudancistrus barbatus  (none) 101000220110010111011000021001
Pseudancistrus brevispinis  (none) ??????????????????????????????
Pseudancistrus sp.  (none) 101000220110010111011000021001
Pseudancistrus sp. Gold Spot  (none) ??????????????????????????????
Pseudolithoxus anthrax  (none) 001000220100010111010000020001
Pseudorinelepis genibarbis  (none) 002000220110010011010000002001
Pterygoplichthys etentaculatus  (none) 101000220100011011010000002001
Pterygoplichthys zuliaensis  (none) 002000220110010011000000002001
Rhinelepis aspera  (none) 102000220100010011110010101001
Rineloricaria rupestris  (none) 000000001000020001000010010100
Schizolecis guentheri  (none) 002000110000000111000000002000
Spectracanthicus murinus  (none) 101000220000010111010000021001
Spectracanthicus punctatissimus  (none) 1010{01}0220000010111010000021
Sturisoma festivum  (none) 000000110110010011000000020101
Sturisomatichthys citurensis  (none) 000000110110010011000000020101
Upsilodus victori  (none) 101000110000010001011010000111

Columns

Column Character Description
1 Anterohyal shape
2 Anterohyal shape
3 Basibranchial 2
4 Basibranchial 3, shape
5 Number of branchiostegals
6 Mesial facing process on branchiostegal 3
7 Length of accessory process of ceratobranchial 1 (ordered)
8 Width of accessory process of ceratobranchial 1 (ordered)
9 Width of ceratobranchial 3
10 Shape of ceratobranchial 5
11 Posteromedial invagination of ceratobranchial 5
12 Connections of ceratobranchial 5
13 Posterolateral process on ceratobranchial 5
14 Accessory process on first epibranchial
15 Mesial surface of epibranchial 1
16 Anterior-facing process on epibranchial 4 located basally to the gill rakers
17 Posterior shelf of epibranchial 4
18 Gill rakers on epibranchial 4
19 Anteromesial projections on hypohyal
20 Width of hypohyal
21 Hypohyal, spindle-shaped
22 Hypobranchial 1
23 Hypobranchial 1
24 Hypobranchial 2
25 Infrapharyngobranchial 4
26 Interhyal
27 Interhyal
28 Interhyal
29 Upper pharyngeal jaw
30 Upper pharyngeal tooth plate
31 Upper pharyngeal tooth plate, shelf lateral to the bulbous section
32 Lateral edge of posterohyal
33 Contact of mesial surface of hyomandibula with quadrate posteroventrally
34 Hyomandibula sutured to pterotic-supracleithrum posterior to cartilaginous condyle of hyomandibula
35 Hyomandibula contacts prootic
36 Anterior margin of hyomandibula sutured to posterior metapterygoid along entire length
37 Opercular condyle of hyomandibula on a process extended beyond posterior margin
38 Length of opercular condyle of hyomandibula
39 Thin, posterior process on hyomandibula just dorsal to opercle
40 Posterior part of hyomandibula beyond opercle
41 Posterior process of hyomandibula incorporated within hyomandibula
42 Posterior region of hyomandibula greatly deflected
43 Ridge on mesial side of hyomandibula located anterodorsally
44 Levator arcus palatini crest (ordered)
45 Levator arcus palatini crest of hyomandibula
46 Hyomandibula deflected beyond posterior margin
47 Process on preoperculo-hyomandibular ridge
48 Preoperculo-hyomandibular ridge continuous
49 Ridge on hyomandibula (contiguous with ridge on quadrate)
50 Zipperlike connection of metapterygoid to lateral ethmoid
51 Anterior connection between metapterygoid and lateral ethmoid
52 Metapterygoid channel
53 Lateral wall of metapterygoid channel
54 Walls of metapterygoid channel
55 Walls of metapterygoid channel
56 Section of metapterygoid dorsal to the anterior process
57 Articulating surface between metapterygoid and lateral ethmoid
58 Anterior process on metapterygoid
59 Ventromesial process of palatine
60 Palatine
61 Orientation of preopercle
62 Exit of preopercular latero-sensory canal
63 Preopercular latero-sensory canal extended posteriorly
64 Quadrate
65 Ventral process on quadrate for articulation with canal plate
66 Quadrate with flap extending below symplectic foramen
67 Articulating condyle of quadrate
68 Longitudinal ridge running the length of the quadrate laterally
69 Angle of dentaries
70 Angle of maxilla
71 Shape of maxilla
72 Premaxilla with cartilaginous connection to mesethmoid
73 Interopercular sesamoid
74 Interoperculo-mandibular ligament
75 Shape of ancistrine opercle (ordered)
76 Double attachment of opercle
77 Maximum forward position of opercle (ordered)
78 Hatchet-shaped opercle
79 Opercle
80 Suprapreopercle
81 Number of rows of plates between suprapreopercle and exposed portion of opercle (ordered)
82 Canal plate
83 Canal plate, number and size
84 Canal plate
85 Contact of canal plate with suspensorium
86 Canal in canal plate
87 Preopercular latero-sensory canal leaves preopercle at first exit and enters a plate
88 Number of cheek plates between the canal plate and opercle (ordered)
89 IO6
90 IO4
91 Number of infraorbitals (ordered)
92 Lateral line
93 Ridge formed by Baudelot?s ligament
94 Contact between frontal and orbit
95 Shape of lateral ethmoid
96 Nasal capsule
97 Ridge on lateral ethmoid
98 Pouch on ventral surface of lateral ethmoid
99 Posterior contact with metapterygoid
100 Mesethmoid disk (ordered)
101 Mesethmoid disk, relative placement
102 Mesethmoid flares anteriorly
103 Mesethmoid anterior edge serrate
104 Mesethmoid continued as a long blade anterior to disk (or well beyond the jaws in species without a disk)
105 Size of nasal
106 Parasphenoid on orbitosphenoid
107 Pterotic-supracleithrum expanded anteroposteriorly
108 Shape of pterotic-supracleithrum
109 Perforations in pterotic-supracleithrum
110 Anterior process of pterotic-supracleithrum
111 Anterior process of pterotic-supracleithrum separated mesially from main body, connected by a strut (ordered)
112 Forward extent of anterior process
113 Bifurcation of anterior process of pteroticsupracleithrum
114 Strut of the pterotic-supracleithrum directed ventrally so that it is visible from below
115 Dorsomesial process on pterotic-supracleithrum
116 Sphenotic
117 Sphenotic, external contact with posteriormost infraorbital
118 Supraoccipital crest
119 Anteriorly directed transverse processes on eighth vertebra
120 Number of vertebrae from first normal neural spine behind dorsal fin to spine under preadipose plate
121 Number of vertebrae from first normal neural spine behind dorsal fin up to, but not including, hypural plate (ordered)
122 Bifid haemal spines
123 Hypurals
124 Posterior margin of the hypural plate
125 First neural spine, positioning
126 Perforations in bifid neural spines
127 Trifid neural spines posterior to dorsal fin
128 Distal margin of the rib of the sixth vertebral centrum
129 Ribs beyond enlarged rib of the sixth vertebral centrum
130 Weberian complex centrum
131 Distal margin of transverse process of Weberian complex centrum
132 Distal margin of transverse process of Weberian complex centrum
133 Tip of transverse process of Weberian complex centrum
134 Transverse process of Weberian complex centrum
135 Tip of transverse processes of Weberian complex centrum
136 Lateral processes of tripus (LPT)
137 Adipose fin
138 Preadipose plates
139 Connecting bone
140 Connecting bone/tendon
141 Connecting bone, contact with nuchal plate
142 Number of dorsal-fin rays
143 Dorsal-fin membrane continues posteriorly
144 Chain-link of proximal dorsal spine to second dorsal-fin pterygiophore
145 Number of dorsal-fin radial elements with transverse processes (ordered)
146 Nuchal plate
147 Nuchal plate
148 Spinelet
149 Spinelet
150 Number of branched anal-fin rays (ordered)
151 First anal-fin pterygiophore with a lateral ridge posterior to the widened anterior surface
152 Adductor fossa
153 Adductor fossa
154 Adductor fossa, exposure (ordered)
155 Shape of cleithrum
156 Shape of posterior margin of the exposed cleithrum
157 Shape of exposed cleithral process
158 Posterior process of coracoid
159 Posterior process of coracoid
160 Posterior section of coracoid
161 Lateral strut of coracoid
162 Lateral strut of coracoid
163 Passage of arrector ventralis through a channel
164 Space between posterior process of coracoid strut and posterior process of coracoid
165 Suture of pectoral girdle
166 Pectoral-fin spines greatly elongated in males
167 Anterolateral processes
168 Anterolateral processes fused to anteromesial processes
169 Anterolateral process of the basipterygium
170 Anteromesial processes
171 Fenestra present anterior to cartilaginous section
172 Ventral ridge of the basipterygium
173 Posteroventral ridge of the basipterygium
174 Posterior process of the basipterygium (Armbruster, 1998b)
175 Lateropterygium of pelvic girdle
176 Two rows of the first pelvic-fin ray
177 First pelvic-fin rays greatly widened in adults
178 Central buccal papilla
179 Buccal papillae
180 Single papilla located behind each dentary
181 Barbel
182 Number of barbels
183 Hypertrophied odontodes on cheeks
184 Evertible cheek plates (ordered)
185 One or two extremely hypertrophied odontodes on cheek
186 Whiskerlike odontodes
187 Fleshy pad covering odontodes on cheeks of nuptial males
188 Hypertrophied odontodes along snout margin anterior to cheek spines
189 Optic notch
190 Position of pectoral fins in relation to first pelvicfin ray when depressed parallel to body axis
191 Number of caudal-fin rays
192 Postdorsal ridge
193 Lips forming suckerlike disk
194 Numerous small plates behind pterotic-supracleithrum
195 Contact between plates prior to anal fin
196 Number of plate rows at thinnest part of caudal peduncle
197 Number of predorsal plates
198 Keeling of plates
199 Hypertrophied odontodes on bodies of nuptial males
200 Extremely elongated odontodes on top of the snout
201 Snout
202 Snout
203 Abdominal plating
204 Teeth in nuptial males
205 Teeth
206 Fleshy appendages around both jaws
207 Lower lip fimbriate
208 Fleshy tentacles on snout (ordered)
209 Tentacules on pectoral fins (ordered)
210 Oesophagus
211 U-shaped diverticulum (ordered)
212 Diverticulum nearly completing a ring
213 Stomach greatly expanded and connected to the abdominal wall by a connective tissue sheet
214 Stomach expanded such that it extends anteriorly to the pectoral girdle and intestine exits dorsally from expanded region
215 Swim bladder