#NEXUS
[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on May 24, 2024; 4:09 GMT

TreeBASE (cc) 1994-2008

Study reference:
Penneys D., & Judd W. 2005. A systematic revision and cladistic analysis of Charianthus
(Melastomataceae) using morphological and molecular characters. Systematic
Botany, 30.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S1346]

BEGIN TAXA;
        TITLE  Combined;
        DIMENSIONS NTAX=14;
        TAXLABELS
            Charianthus_alpinus
            Charianthus_corymbosus
            Charianthus_dominicensis
            Charianthus_grenadensis
            Charianthus_nodosus
            Charianthus_purpureus
            Clidemia_guadalupensis
            Graffenrieda_latifolia
            Leandra_lima
            Miconia_howardiana
            Miconia_laevigata
            Tetrazygia_bicolor
            Tetrazygia_crotonifolia
            Tetrazygia_fadyenii
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M1415]

	TITLE Combined;
	LINK TAXA = Combined;
	DIMENSIONS NCHAR=776;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 a t c g r y k m s w n J K M N P Q R S T U V W X Y Z 7 b 9 d e f A h j D E H p q B F 8 u v G x C" MISSING=? GAP= -;

MATRIX
[                                    10        20        30        40        50        60        70        80        90        100       110       120       130       140       150       160       170       180       190       200       210       220       230       240       250       260       270       280       290       300       310       320       330       340       350       360       370       380       390       400       410       420       430       440       450       460       470       480       490       500       510       520       530       540       550       560       570       580       590       600       610       620       630       640       650       660       670       680       690       700       710       720       730       740       750       760       770       780  ]
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Charianthus_alpinus          GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCMCAGCTAACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGACGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACCTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCCTCGATAGTCCTCGACGCGGGACGGGGG-------ATGCACGCAA-TCTCTACTCGA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTCGAGTATATACCCGAGCGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG100000?1001000000100000{01}100002110330001000101??002000331?221100010101110
Charianthus_corymbosus       GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCACAGCTAACTGGACTGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGACGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCCTTGATAGTCCTCGACGCGGGACGGGGG-------ATGCACGCAA-TCTCTACTCGA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTCGAGTATATACCCGAGCGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG100000?100100000000000011000021{12}0030001001101??001000231?221100010001110
Charianthus_dominicensis     GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCACAGCTAACTGGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGYCGCGTCGACGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCCTTGATAGTCCTCGACGCGGGACGGGGG-------ATGCACGCAA-TCTCTACTCGA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTCGAGTATATACCCGAGCGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG100001010{01}001000100100011000021{12}0030001001101??001000231?221100010001110
Charianthus_grenadensis      GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCACAGCTAACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCG?????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????????CGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGC???????????????????????????????????????????????????????????????????????????????????????????1000011100100000100000011000021{12}0330001000101??002000331?221100210101110
Charianthus_nodosus          GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCACAGCTAACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGACGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCCTTGATAGTCCTCGACGCGGGACGGGGG-------ATGCACGCGA-TCTCTACTCGA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTCGAGTATATACCCGAGCGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG100001210101000011000114100002110330001000101??002000331?221211110001110
Charianthus_purpureus        GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCACAGCTAACTGGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGACGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCCTTGATAGTCCTCrACGCG??????????????????????????????????????????????????????CGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TGTATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGACGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTCGAGTATATACCCGAGCGCGATCCCAGGTCAGGCGGG???????????????????100001210{01}010000100{01}0001100002110330001000101??002000331?221100010001110
Clidemia_guadalupensis       GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCATTGCGTACTAGACAGAGCCCAAGAGCACCTAACGCTCATGGGGCTCGCGTCGTGTCGGCGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAATGTCAGCACGGATCGTGCCAAGGAGCTTTATTGAGAGAATCGTCGTCCCACGCCTTTGATAGTCCTCGATGCGCGACGGGGGGCGGGGGACGTACGCAA-TCTCTACCCAA-AGTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGCGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCTTACGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGG---TAT-TATGCTCGGGCGCAGAATGTGGTCTCCCGTTGGACTGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CTGGACTTTGAATAT--CCCCGAGTGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTtTAAG1000112?0200000000000323100000000210001100011??00010013001211302100?1110
Graffenrieda_latifolia       GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAACCGTCACCGCGTACTAGAGAGACCTCGTGAGCACCCAAAGCGCACGCGGCTCGCGTCGCGTCGGCGCGGGAGT-ACTAG-AGACCCGCGTCGTACCGGCGAACGAACAACTTCAGCACGGATCGTGCCAAGGAGCTGCATTTTGAGAATCGTCGTCCCACGCCCCC-ATCGTCCTCGATGCGGGACGGGGG-------ATGCACGCAA-TCTCTGATAAG--GTCTGAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGCGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGTCACCC-GCACGAGCATCGGCGGGGATTATATACGCGAGGGCGCAGAATATGGTCTCCCGTGGGATCGGCATGCGCGGCATGCCGAGGCGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTCGGGTGGGATCGAACGAGATTGGCCGCGTCGTGGTCTCGGCATGCTGCGGGCT-CCGGACTTTGGCAAT--CCCCGAGTACGATCCCAGGTCAGGCGgggATACCCGCTGAGTtTAAG000000??0000000000000000000000000000000?00000000000000000000000000010000
Leandra_lima                 GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCATTGCTCACTAGACAGAGCCCGTGAGCGCCTAGCGCTCATGGGGCTCGCGTCGCGTCGGCGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGTCCTCGAT----------GCGGGACGGGGG-------ATGTACGCAA-TCTCTACTCAA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGCGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA-TAT--ATGCTGGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGACGAGGTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGA?ATTGGCTGC-CCGTGGGCTCGGCATGCGGCGG-CTTCCGGACTTTGAATAT--?CCCGAGTGCGATCCCA?GTCA?GCGGGGCTACCCGTTGAGTTTAAG1001012?030001002201020210011000212210101101001000410131?2211201000?1110
Miconia_howardiana           GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCATTGCGTACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGACTCGCGTCGCGTCGGCGTGGGTGT-ACTCG-AGACCAGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCATGGAGAGAATCGTCGTCCCACGCCTTTGATAGTCCTCGATGCGGGACGGGGGGCGGGGGATGTAC-CA--TCTCTACTCAA--GTCTAAaCGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGCGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TATATGCTCGGGCGCAGAATATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGCAATGGCTGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATTAGGCGGGCT-CCGGACTTTGAGTAT--CCCCGAGTACGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTtTAAG1001012?00000001300100021100120{13}0000001010101??0012201102231??0201011110
Miconia_laevigata            GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCGTTGCGTACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGGCGCGGGTGTGCACTGGAGACCCGCGTCGTACCGGCGGACAAACAATGTCAGCACGGATCGTGCCAAGGAGCTTTATGGAGAGAATCGTCGTCCCACGCCTTTGGTGGTCCTCGATGCGGGACGGGGGGCGAGGGATGTACGCAA-TCTCTACTCGA--GTCAAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGCGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCTTGGGCACCCCGCATGAGCATTGGCGGGGA---TATACGCTCGGGCGCAGAATATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGTCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTTGAATAT--CCCCGAGTGCGATCCCAGGTCAGGCGGGGCTAC?CGCTGAGTTTAA?1000014?00000000010100021000110{03}0120001010000020100001100111000101011110
Tetrazygia_bicolor           GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAACCGTCATTGCTTACTAGACAGAGCCCGTGAGCGCCTAACGCTCGTGGGGCTCGCGCCGCGTCGGCGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCAAGGAGAGAATCGTCGTCCCACGCCTTTGATAGTCCTCGACGCGGGACGGGGG-------ACGTACGCAAATCTCTACTCAAAAGTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCCCGCATGAGCATTGGCGGGGA-TATATATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCACGTCGAGCTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGAGCACCGCGGCGTTCGGTGG-ATCGAATCAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTTGAATAT--CCCCGAGTGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG101000??1000000002001000101012001001011010021??10000022002111001000?1110
Tetrazygia_crotonifolia      GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCATTGCTTACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGTCGCGTCGGCGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCTGCACGGAGAGAATCGTCGTCCCACGCCTTTGATAGTCCTCGACGCGGGACGGGGG-------ACGTACGCAA-TCTCTACTCAA--GTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCACGAGCATTGGC--------------CTCGGGCGCAGAACATGGTCTCCCGCGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATGAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTTGAATAT--CCCCGAGAACGATCCCAGGTCAGGCGGGACTACCCGCTGAGTTTAAG110101301000001002110000100001001000011002011??10100113002211002000?1111
Tetrazygia_fadyenii          GCGGAAGGATCATTGTCGAAATCGAACGGAAGAACGACCCGCGAATCGTCATTGCTTACTAGACAGAGCCCGTGAGCACCTAACGCTCATGGGGCTCGCGCCGCGTCGGCGCGGGTGT-CCTCG-AGACCCGCGTCGTACCGGCGGACAAACAACGTCAGCACGGATCGTGCCAAGGAGCCGCAAGGAGAGAATCGTCGTCCCACGCCTTTGATAGTCCTCGACGCGGGACGGGGG-------ACGTACGCAA-TCTCTACTCAAAAGTCTAAACGACTCTCGGCAACGGATATCTCGGCTCTTGCATCGATGAAGAACGTAGCGAAATGCGATACTTGGTGTGAATTGCAGAATCCCGTGAACCATCAATTCTTTGAACGCAAGTTGCGCCCGAAGCCATTCGGCCGAGGGCACGCCTGCCTGGGTGTCGTGAATCCCCTTGCCCCTAGGCACCC-GCATGAGCATTGGCGGGGA---TATATGCTCGGGCGCAGAACATGGTCTCCCGTGGGATCGGCATGCACGGCATGTCGAGGTGCGGTTGGCCGAAAATCGAGC-ATGGCGGTGACGGGCACCACGGCGTTCGGTGG-ATCGAATCAGATTGGCTGCGCCGTGGGCTCGGCATGCGGCGGGCT-CCGGACTTTGAATAT--CCCCGAGTGCGATCCCAGGTCAGGCGGGGCTACCCGCTGAGTTTAAG100000?000100000020010001000000113010110000201?1010003301201100001001110
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