#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on April 26, 2024; 14:24 GMT TreeBASE (cc) 1994-2008 Study reference: Zhang X., Smith M.R., Yang J., & Hou J. 2016. Onychophoran-like musculature in a phosphatized Cambrian lobopodian. Biology Letters, . TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S18871] BEGIN TAXA; TITLE Taxa1; DIMENSIONS NTAX=48; TAXLABELS Acinocrinus Actinarctus Aegirocassis_benmoulae Alalcomenaeus Anomalocaris_canadensis Antennacanthopodia Aysheaia Carbotubulus Cardiodictyon Chengjiangocaris Collins_monster_Burgess_Shale Collins_monster_Emu_Bay Collinsium Cricocosmia Diania Euperipatoides_Onychophora Fuxianhuia Hadranax Hallucigenia_fortis Hallucigenia_hongmeia Hallucigenia_sparsa Halobiotus Helenodora Hurdia_victoria Jianshanopodia Kerygmachela Kuamaia_lata Leanchoilia Luolishania Lyrarapax_unguispinus Megadictyon Microdictyon Misszhouia_longicaudata Onychodictyon_ferox Onychodictyon_gracilis Ooperipatellus Opabinia Orstenotubulus Pambdelurion Paucipodia Peytoia_nathorsti Plicatoperipatus_Onychophora Schinderhannes Siberian_Orsten_tardigrade Siberion Tritonychus Tubiluchus Xenusion ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M37231] TITLE Panarthropods; LINK TAXA = Taxa1; DIMENSIONS NCHAR=115; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5" MISSING=? GAP= -; CHARSTATELABELS 1 Paired_appendages / absent present, 2 Anterior_region_covered_by_sclerites / absent present, 3 'Head shield (cephalic shield) formed by fused cephalic segments' / absent present, 4 'Isolated dorsal sclerite associated with eye-stalks' / isolated_dorsal_sclerite_absent_or_not_associated_with_eyestalks isolated_dorsal_sclerite_is_associated_with_eyestalks, 5 Shape_of_dorsal_isolated_sclerite / oval_or_rounded elongate triangular, 6 Degree_of_attachment_of_dorsal_isolated_sclerite_on_head / broad_attachment_to_cephalic_region narrow_attachment_to_anterior_edge_of_cephalic_region, 7 'Isolated lateral sclerites, forming tripartite carapace' / absent present, 8 Anterior_trunk_flexure_in_coronal_plane / orientation_of_mouth_is_fixed_relative_to_main_trunk 'flexible anterior trunk allowing mouth’s dorsal-ventral orientation to be independent of main trunk axis', 9 Swelling_of_anteriormost_trunk / 'anteriormost trunk contiguous with posterior trunk; no swollen ‘head’' 'anteriormost trunk elliptical, substantially wider than adjacent trunk', 10 'Spine-like cuticular projections on anterior region (cirri)' / absent present, 11 Mouth_opening_orientation / terminal ventral posterior, 12 Appendages_anterior_to_mouth_opening / no_appendages_anterior_to_mouth_opening one_or_more_appendage_pairs_anterior_to_mouth_opening, 13 'Pre-oral chamber' / absent present, 14 Radially_arranged_circumoral_structures / absent present, 15 Differentiated_circumoral_structures / 'undifferentiated plates (e.g. Pambdelurion)' 'differentiation of three or four enlarged plates (i.e. Radiodonta)', 16 Spinose_projections_from_inner_face_of_circumoral_structures / proximal_surface_with_single_projection proximal_surface_with_multiple_spines, 17 Pharynx_differentiated_from_midgut / not_differentiated differentiated, 18 Pharynx_eversible / permanently_inverted completely_eversible, 19 Sclerotized_pharyngeal_‘teeth’ / absent present, 20 Nature_of_pharyngeal_teeth_or_aciculae / 'spinose/acicular: each tooth has a single point' 'multiple cusps: each tooth has multiple tips, perhaps expressed as denticles or serrations', 21 Arrangement_of_pharyngeal_teeth_or_aciculae / uniform_distribution_around_pharynx limited_number_of_longitudinal_rows_or_series, 22 Eyes / absent present, 23 Eye_attachment / eye_sessile eye_stalked, 24 Type_of_eyes / 'ocellus-like or pigment spots ' 'multiple visual units (including compound eyes) ', 25 'Sclerotized post-ocular (post-protocerebral) body appendages with arthrodial membranes' / 'not sclerotized; arthrodial membranes absent (‘lobopodous’)' 'sclerotized; arthrodial membranes present (‘arthropodized’)', 26 'Pre-ocular (protocerebral) limb pair, structurally differentiated from trunk appendages' / 'pre-ocular limb pair absent or not differentiated from other limbs' 'distinct pre-ocular limb pair', 27 'Sclerotization of pre-ocular (protocerebral) limb pair ' / not_sclerotized sclerotized, 28 'Pre-ocular (protocerebral) limb pair with arthrodial membranes ' / absent present, 29 'Nature of post-ocular lobopodous inner branch ' / 'cylindrical/subconical appendage' laterally_expanded_swimming_flap, 30 Deutocerebral_limb_pair_structurally_differentiated_from_trunk_appendages / 'undifferentiated, or differentiated in size only' structurally_differentiated, 31 'Nature of sclerotized first post-ocular (deutocerebral) appendage' / antenniform_with_distinct_podomeres 'short great-appendage', 32 'Nature of lobopodous first post-ocular (deutocerebral) appendage' / ambulatory sensorial 'masticatory, with sclerotized jaw', 33 Inner_blade_of_deutocerebral_jaw_with_diastema / absent present, 34 'Nature of lobopodous second post-ocular (tritocerebral) appendage' / undifferentiated specialized_papilla, 35 'Nature of arthropodized second post-ocular (tritocerebral) appendage' / ambulatory_limb_with_distinct_podomeres 'specialized post-antennal appendage', 36 'Position of pre-ocular (protocerebral) appendage pair' / lateral ventral terminal, 37 'Pre-ocular (protocerebral) appendages directly adjacent to one another' / 'pre-ocular appendages not directly adjacent' 'pre-ocular appendages adjacent to one another, with or without physical fusion', 38 'Pre-ocular (protocerebral) appendages mechanically fused' / 'pre-ocular appendages adjacent but not mechanically fused' 'pre-ocular appendages are mechanically fused to form a single element', 39 'Extent of pre-ocular (protocerebral) appendage fusion' / 'locational basal only, with separate distal elements ' fused_into_a_reduced_labrum_, 40 'Spines/spinules on pre-ocular (protocerebral) appendage' / absent 'present (anomalocaridids, gilled lobopodians, certain lobopodians)', 41 'Number of spine/spinule series on pre-ocular (protocerebral) frontal appendage' / 'one series (e.g. Aysheaia, Kerygmachela, Opabinia, Hurdia, Peytoia)' 'two series (e.g. Anomalocaris, Onychodictyon ferox)', 42 'Coplanar spine/spinule series in pre-ocular (protocerebral) frontal appendages' / 'no (as in Radiodonta)' 'yes (as in Onychodictyon ferox)', 43 Multifurcate_distal_termination_of_protocerebral_appendage / absent present, 44 Epidermal_segmentation / absent present, 45 Dorsal_integument_sclerotized_and_connected_by_arthrodial_membranes / absent present, 46 Sternites_connected_by_arthrodial_membranes / absent_ present, 47 Annulations / absent present, 48 Annulation_distribution / limbs_only_ trunk_and_limbs_, 49 Anterior_projection_of_trunk_lacking_annulations / annulations_continue_for_full_length_of_trunk differentiated_anterior_region_of_trunk_lacking_annulations, 50 Organization_of_trunk_annulation / homonomous heteronomous, 51 Branching_of_annular_rings / unbranched branched, 52 Metamerically_arranged_dorsolateral_epidermal_specializations / absent present, 53 Nature_of_paired_epidermal_specialization / epidermal_depressions epidermal_evaginations, 54 Proportions_of_epidermal_trunk_evaginations / 'wider than tall (e.g. nodes or plates)' 'taller than wide (e.g. spines)', 55 Trunk_epidermal_evaginations_with_acute_distal_termination / absent present, 56 Acute_distal_termination_in_epidermal_evagination_is_curved / straight curved, 57 Sclerotization_of_epidermal_evaginations / epidermal_evaginations_not_sclerotized epidermal_evaginations_sclerotized, 58 Dorsal_trunk_sclerite_ornament / 'net-like' scaly, 59 Sclerites_consist_of_a_stack_of_constituent_elements / sclerites_comprise_single_element sclerites_comprise_stacked_elements, 60 Maximum_number_of_primary_dorsal_epidermal_specializations_above_each_leg_pair / one two three four five seven, 61 ‘Secondary’_structures_in_dorsal_epidermal_specializations / absent present, 62 Size_of_dorsal_epidermal_specialisations_consistent / each_group_of_dorsal_elements_of_equivalent_size size_of_dorsal_elements_varies_between_groups, 63 Spacing_of_epidermal_specializations / regular variable, 64 Papillae_on_trunk_annulations / absent_ present, 65 'Serially repeated mid-gut glands' / absent 'reniform, submillimetric lamellar', 66 Differentiated_anterior_trunk / trunk_of_uniform_construction 'anterior trunk differentiated from posterior trunk by abrupt change in thickness, armature and appendage construction', 67 Trunk_exites / absent present, 68 Form_of_exite / lateral_lobes simple_oval_paddle_with_marginal_spines bipartite_shaft_with_lamellar_setae_, 69 'Dorsal flaps / exites fused with endopod to form biramous appendage' / not_fused fused, 70 'Exites (= lanceolate dorsal blades) associated with dorsolateral flaps' / absent present, 71 'Exite (=setal blade) distribution' / confined_laterally present_dorsally, 72 'Antero-posteriorly compressed protopodite with gnathobasic endites in post-deutocerebral appendage pair' / absent present, 73 'Secondary structures on non-sclerotized (lobopodous) limbs' / absent present, 74 Nature_of_secondary_structure / 'spines/setae' appendicules, 75 Length_of_spines_on_secondary_structure / 'short / equant' 'needle-like', 76 'Papillae on non-sclerotized (lobopodous) limbs' / absent present, 77 Papillae_with_terminal_spine / spine_absent spine_present, 78 'Finger-like elements in distal tip of limbs' / absent present, 79 Terminal_claws_on_trunk_limbs / absent present, 80 Terminal_claws_with_multiple_branches / absent present, 81 Number_of_claws_on_undifferentiated_trunk_limbs / one two three four seven, 82 Nature_of_claws_on_each_trunk_limb / claws_on_single_limb_all_identical_ claws_on_single_limb_differentiated, 83 Differentiated_distal_foot_in_lobopodous_trunk_limbs / absent present, 84 Strengthening_rays_in_lateral_flaps / absent present, 85 Posterior_tapering_of_lateral_flaps / absent present, 86 Hypertrophied_set_of_anterior_body_flaps / absent present, 87 Anterior_flaps_reduced / anterior_flaps_present anterior_flaps_reduced_in_size, 88 Number_of_lobopodous_limbs_on_differentiated_anterior_trunk / two three five six, 89 Nature_of_lobopodous_limbs_on_differentiated_anterior_trunk / 'slender, simple' cirrate, 90 Appendages_comprise_15_or_more_podomeres / Fewer_than_15_podomeres 15_or_more_podomeres, 91 Limbless_posterior_extension_of_the_lobopodous_trunk / absent 'present: tubular portion of the body extends beyond the last observable appendage pair', 92 Posterior_tagma_composed_of_three_paired_lateral_flaps / absent present, 93 Posteriormost_pair_of_trunk_appendages_structurally_differentiated / undifferentiated differentiated, 94 Nature_of_differentiated_posteriormost_appendages / appendicular_tail 'partially fused/reduced walking legs', 95 Nature_of_appendicular_tail / tail_rami tail_flaps, 96 Direction_of_claws_on_posteriormost_appendage_pair / Same_direction_as_claws_on_other_appendages rotated_anteriad, 97 Ventral_nerve_cord_with_paired_ganglia / absent present, 98 Dorsal_condensed_brain / absent present, 99 Number_of_neuromeres_integrated_into_the_dorsal_condensed_brain / one_ two three_, 100 Mouth_innervation_relative_to_brain_neuromeres / protocerebral_innervation deutocerebral_innervation innervation_from_multiple_neuromeres tritocerebral_innervation, 101 'Ventral nerve cord (VNC) paired' / unpaired paired, 102 VNC_with_morphologically_discrete_condensed_hemiganglia_connected_by_medial_commissures / hemiganglia_absent morphologically_discrete_condensed_hemiganglia_connected_by_medial_commissures, 103 Paired_nerve_cord_lateralized / 'absent (Alalcomenaeus, Fuxianhuia, Tardigrada)' 'present (Onychophora)', 104 Paired_nerve_cord_bears_medial_interpedal_commissures / medial_interpedal_commissures_absent medial_interpedal_commissures_present, 105 Regularly_spaced_peripheral_nerves_along_the_entire_length_of_the_nerve_cord / 'absent, or not occurring regularly along entire length of nerve cord' present_along_entire_length_of_nerve_cord, 106 Nerve_cord_has_orthogonal_organization / not_orthogonally_organized orthogonally_organized, 107 Orthogonal_nerve_cord_has_complete_ring_commissures / ring_commissures_incomplete_or_absent complete_ring_commissures, 108 Segmental_leg_nerves_shifted_anteriorly_relative_to_appendages / not_shifted_anteriorly shifted_anteriorly, 109 Segmental_leg_nerves_paired / unpaired paired, 110 Stomatogastric_ganglion / absent present, 111 Heart / absent present, 112 Skeletal_musculature / peripheral_longitudinal_and_circular_muscle metamerically_arranged_skeletal_muscle, 113 Longitudinal_peripheral_musculature / absent present, 114 Circular_peripheral_musculature / absent present, 115 Circular_musculature_inside_longitudinal_musculature / circular_muscles_inside_longitudinal longitudinal_muscles_inside_circular, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 ] [ . . . . . . . . . . . ] Acinocrinus 1???????????????????????0???00-{01}-0-????????00-11?10111111??511??010-----101????????----21-?-??????????????????????? Actinarctus 1100---00110010010100100011000-0-0-10------00-0----1100-100??---000-----0--0-110310-------0-11-11100110111011?-100- Aegirocassis_benmoulae 11011010?0?10???????????011110-0-0-1???10-100-0----0------------?010011-0--0-00----110?---000---??????????????????? Alalcomenaeus 1110---0--2100--?0???11111??-11---01111----11?------------0-----1-111-01-----0100--------0--0---112111000--???????? Anomalocaris_canadensis 11010000?011011110???111011110-0-0-110-110100-0----0------------1010001-0--0-00----1101---01101-???????????????1??? Antennacanthopodia 10-----?00???????????100010-01-1-0-00--0---00-1????0------?----??00-----0--1?00-----------1-0---??????????????????? Aysheaia 10-----0000-0100?0???0--010-00-0-0-00--10-100-111000------0----1000-----1000-010400-------0-11-1???????????????0??? Carbotubulus 1???????????????????????0???0?-???-????????00-0----????????????-?10-----0--0-??????----??-0-??-????????????????0??? Cardiodictyon 10-----110{01}??????0???10?0???0?-??0-00--0---00-111101100-10?1-000000-----0--0-0101?0-------1-0--0??????????????????? Chengjiangocaris 110101-0--2100--?0???1111110-10---11111----110------------------0-111-00-----00----------1--101-1?????0?1?????????? Collins_monster_Burgess_Shale 1???????????????????????0???00-{01}-0-????????00-11?10111111????????10-----101????????----31-?-??????????????????????? Collins_monster_Emu_Bay 110?????????????????????0???0?-{01}-0-????????00-11?10111111??201???10-----1010-0100-0----21-????????????????????????? Collinsium 1100---1000-?????????0--010-00-{01}-0-00--0---00-110101111110140111010-----1010-0100-0----31-1-0--0??????????????????? Cricocosmia 00-----0000-0100111?00---------------------00-1-1001100-1001-00100------------------------------???????????????0??? Diania 10-----0000-?????0???0--00--00-0-0-00--0---00-110101100-0??0-001000-----1000-0?---0-------1-0--???????????????????? Euperipatoides_Onychophora 10-----0001110--100--100010-01-201-00--0---00-110010------1----1000-----0--11010101-------1-0--00112101111101010111 Fuxianhuia 110101-0--2100--?0???1111110-10---11111----110------------------0-111-00-----00----------1--101-?12111????????1100- Hadranax 1???????????????????????0???0?-???-????????00-11?101100-0-?3--01?00-----0--10?????0-------?-??????????????????????? Hallucigenia_fortis 10-----1100-?????0???10?00--00-{01}-0-00--0---00-1111?111111??10000010-----0--0-0101?0----00-?-0--0??????????????????? Hallucigenia_hongmeia 1???????????????????????0???0?-??0-????????00-11?1?1111110?10100??0-----0--0-0100-0----??-1-0--???????????????????? Hallucigenia_sparsa 10-----1100-11001010110000--00-{01}-0-00--0---00-0----111111111000-010-----0--0-010100----10-0-0--0??????????????????? Halobiotus 10-----0000-010010100100011000-0-0-20------00-0----10-----0??---000-----0--0-111110-------0-11-111001101110111-100- Helenodora 10-----?00??????????????010-0?-??1-00--0---00-?????0------?----10?0-----0--1?01010?----??-?-??????????????????????? Hurdia_victoria 11011010?011011110110111011110-0-0-1???10-1?0-0----0------------?010011-0--0-00----1001---00101???????????????????? Jianshanopodia 10-----0000-01??101100--010-00-0-0-0???10-?00-11?100------?----0100-----11-0-0????0-------0-101-??????????????????? Kerygmachela 10-----0000-0100?0??????010-00-0-0-010-10-100-110101100-0-?3-?011010010-0--0-00---00000---00100-???????????????0??? Kuamaia_lata 111101-0--2100--?0???1111110-10---01111----111------------0-----1-121-01-----01021-------0--0---???????????????1??? Leanchoilia 1110---0--2100--?0???1111110-11---01111----11?------------0-----1-111-01-----010{02}?-------0--0---??????????????????? Luolishania 1100---?100-?????0???100010-00-{01}-0-00--0---00-11110111111??20111010-----101100100-0----21-1-0--0??????????????????? Lyrarapax_unguispinus 11010000?01101???????111011110-0-0-110-10-100-0----0------------0010001-0--0-00----?111---01101-?10011?????????1??? Megadictyon 10-----?00???1???0???0--010-00-0-0-00--10-100-11?0?0------?----0100-----11-0-0????0-------?-??????????????????????? Microdictyon 10-----1000-?????0???0--00--00-0-0-00--0---00-111101100-10?1-000000-----0--0-0101?0-------1-0--0??????????????????? Misszhouia_longicaudata 111101-0--2100--?0???1111110-10---01111----111------------0-----1-121-01-----0100--------0--0---???????????????1??? Onychodictyon_ferox 10-----0000-0????0???100010-00-0-0-00--111000-111101101010?1-001000-----11-0-010110-------0-11-1??????????????????? Onychodictyon_gracilis 10-----??00-0????0??????0???0?-?-0-????????00-110?0110??1??1-0010?0-----???0-0101?0----??-1-11-0??????????????????? Ooperipatellus 10-----0001110--100--100010-01-211-00--0---00-110010------1----1000-----0--11010101-------1-0--00112101111101010111 Opabinia 10-----000210????0???111010-00-0-0-111010-110-0----0------------1010011-0--0-00---00000---?1101-??????????????????? Orstenotubulus 1???????????????????????0???0?-???-????????00-11?11111?????101?1??0-----0--11??????----??-?-??????????????????????? Pambdelurion 10-----000100100?0???0--010-00-0-0-110-10-?00-1????????????--?-?1010010-0--0-00---00000---?????????????????????110- Paucipodia 10-----0000-?????01??0--00--00-0-0-00--0---00-110000------?----0000-----0--0-010100-------1-0--???????????????????? Peytoia_nathorsti 11011000?0110111?0???111011110-0-0-110-10-100-0----0------------1010011-0--0-00----1101---000---??????????????????? Plicatoperipatus_Onychophora 10-----0001110--100--100010-01-211-00--0---00-110010------1----1000-----0--11010101-------1-0--00112101111101010111 Schinderhannes 1??????0?01101???????1110111??-0-0-110-110?????????????????????????????-?????????????1?---00101-??????????????????? Siberian_Orsten_tardigrade 10-----0000-0100????????0???0?-???-????????00-0----???--?????---?00-----0--0-011110-------0-1{12}-???????????????????? Siberion 10-----0000-0100?0???0--010-00-0-0-010-????00-11?000------?----1?00-----???0-??????-------?-??????????????????????? Tritonychus 1???????????????????????0???0?-???-????????00-11??1????????????1??0-----0--1101?200----????????????????????????01?? Tubiluchus 00-----0000-0100111100---------------------00-1-1000------0----100------------------------------00--00--111--000111 Xenusion 10-----?00???????????0--00--00-0-0-00--????00-11?101100-0??1-001?00-----1??????????-------1-0--???????????????????? ; END; BEGIN TREES; TITLE Panarthropods_Consensus; LINK TAXA = Taxa1; TRANSLATE 1 Tubiluchus, 2 Cricocosmia, 3 Aysheaia, 4 Siberion, 5 Onychodictyon_ferox, 6 Onychodictyon_gracilis, 7 Diania, 8 Xenusion, 9 Paucipodia, 10 Microdictyon, 11 Cardiodictyon, 12 Hallucigenia_sparsa, 13 Hallucigenia_fortis, 14 Hallucigenia_hongmeia, 15 Luolishania, 16 Collinsium, 17 Collins_monster_Burgess_Shale, 18 Collins_monster_Emu_Bay, 19 Acinocrinus, 20 Orstenotubulus, 21 Tritonychus, 22 Carbotubulus, 23 Antennacanthopodia, 24 Helenodora, 25 Euperipatoides_Onychophora, 26 Plicatoperipatus_Onychophora, 27 Ooperipatellus, 28 Actinarctus, 29 Halobiotus, 30 Siberian_Orsten_tardigrade, 31 Megadictyon, 32 Jianshanopodia, 33 Hadranax, 34 Kerygmachela, 35 Pambdelurion, 36 Opabinia, 37 Anomalocaris_canadensis, 38 Peytoia_nathorsti, 39 Hurdia_victoria, 40 Aegirocassis_benmoulae, 41 Lyrarapax_unguispinus, 42 Schinderhannes, 43 Fuxianhuia, 44 Chengjiangocaris, 45 Leanchoilia, 46 Alalcomenaeus, 47 Misszhouia_longicaudata, 48 Kuamaia_lata; TREE 'Majority-Rule' = [&R] (1,(2,(3,((6,((20,21,23,24,25,26,27),((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))))),((5,(28,29,30)),(33,((31,32),(4,(34,(35,((38,39,40,(37,(41,42))),(36,((43,44),((45,46),(47,48))))))))))))))); END; BEGIN TREES; TITLE Panarthropods_Results; LINK TAXA = Taxa1; TRANSLATE 1 Tubiluchus, 2 Cricocosmia, 3 Aysheaia, 4 Siberion, 5 Onychodictyon_ferox, 6 Onychodictyon_gracilis, 7 Diania, 8 Xenusion, 9 Paucipodia, 10 Microdictyon, 11 Cardiodictyon, 12 Hallucigenia_sparsa, 13 Hallucigenia_fortis, 14 Hallucigenia_hongmeia, 15 Luolishania, 16 Collinsium, 17 Collins_monster_Burgess_Shale, 18 Collins_monster_Emu_Bay, 19 Acinocrinus, 20 Orstenotubulus, 21 Tritonychus, 22 Carbotubulus, 23 Antennacanthopodia, 24 Helenodora, 25 Euperipatoides_Onychophora, 26 Plicatoperipatus_Onychophora, 27 Ooperipatellus, 28 Actinarctus, 29 Halobiotus, 30 Siberian_Orsten_tardigrade, 31 Megadictyon, 32 Jianshanopodia, 33 Hadranax, 34 Kerygmachela, 35 Pambdelurion, 36 Opabinia, 37 Anomalocaris_canadensis, 38 Peytoia_nathorsti, 39 Hurdia_victoria, 40 Aegirocassis_benmoulae, 41 Lyrarapax_unguispinus, 42 Schinderhannes, 43 Fuxianhuia, 44 Chengjiangocaris, 45 Leanchoilia, 46 Alalcomenaeus, 47 Misszhouia_longicaudata, 48 Kuamaia_lata; TREE piwe_k_93.12 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_1.061 = [&R] (1,(2,(3,((5,((4,33,34,(35,((28,(29,30)),((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_65.84 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_40.74 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_31.09 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_28.71 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_equal = [&R] (1,(2,(3,4,5,6,7,8,9,10,11,13,14,20,21,23,24,25,26,27,33,34,35,(12,22),(15,16,17,18,19),(28,29,30),(31,32),((36,((43,44),((45,46),(47,48)))),(37,38,39,40,41,42))))); TREE xpiwe_k_259.4 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_65.84 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(30,(28,29)))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_259.4 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_57.29 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_117.3 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_117.3 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_45.19 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_37.01 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_159.5 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_57.29 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_45.19 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_40.74 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_37.01 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_33.83 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_159.5 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_93.12 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_77.20 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),((37,(41,42)),(38,(39,40))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_77.20 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(39,(40,(38,(37,(41,42))))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_50.59 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(30,(28,29)))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_50.59 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_33.83 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE xpiwe_k_31.09 = [&R] (1,(2,(3,(((33,((4,(34,(35,((36,((43,44),((45,46),(47,48)))),(38,39,40,(37,(41,42))))))),(31,32))),(5,(28,29,30))),(6,(((7,8),(9,(10,(11,(13,(12,22),(14,(15,16,17,18,19))))))),(20,21,23,24,25,26,27))))))); TREE piwe_k_equal = [&R] (1,(2,(3,4,5,6,7,8,9,10,11,13,14,20,21,23,24,25,26,27,33,34,35,(12,22),(15,16,17,18,19),(28,29,30),(31,32),((36,((43,44),((45,46),(47,48)))),(37,38,39,40,41,42))))); END;