#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on April 16, 2024; 23:53 GMT TreeBASE (cc) 1994-2008 Study reference: Kavanagh F., & Wilson G.D. 2007. A revision of the genus Haplomesus (Isopoda: Asellota: Ischnomesidae) with erection of four new genera. Invertebrate Systematics, 21: 1-49. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S1924] BEGIN TAXA; TITLE Taxa1; DIMENSIONS NTAX=39; TAXLABELS Contrarimesus_curtispinis Contrarimesus_franklinae Cornuamesus_biscayensis Cornuamesus_longiramus Fortimesus_brevispinis Fortimesus_concinnus Fortimesus_consanguineus Fortimesus_cornutus Fortimesus_gigas Fortimesus_profundicolus Fortimesus_robustus Fortimesus_scabriusculus Fortimesus_thomsoni Fortimesus_zuluensis Gracilimesus_angustus Gracilimesus_celticensis Gracilimesus_corniculatus Gracilimesus_gorbunovi Gracilimesus_gorbunovi_sensu_Svavarsson Gracilimesus_hanseni Gracilimesus_insignis Gracilimesus_orientalis Gracilimesus_tenuispinis Gracilimesus_tropicalis Haplomesus_bifurcatus Haplomesus_ornatus Haplomesus_quadrispinosus Haplomesus_quadrispinosus_sensu_Brandt Heteromesus_drachi Heteromesus_frigidus Heteromesus_granulatus Heteromesus_schmidtii Heteromesus_similis Heteromesus_spinosus Ischnomesus_antarcticus Ischnomesus_bispinosus Stylomesus_hexapodus Stylomesus_menziesi Stylomesus_sarsi ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2925] TITLE Ischnomesidae_morphlogical; LINK TAXA = Taxa1; DIMENSIONS NCHAR=79; FORMAT DATATYPE=Standard SYMBOLS= "1 2 3 4 5" MISSING=? GAP= -; CHARSTATELABELS 1 Per.5_fusion / freely_articulated fused_with_pereonite_6, 2 Per.6_fusion / freely_articulated fused_with_pereonite_7, 3 Per.7_fusion / freely_articulated fused_with_pleonite_1, 4 PL_1_fusion / freely_articulated fused_with_pleotelson, 5 Haplomesus_segmental_artic. / 'articulation absent- no segmental demarcation' 'articulation present- rudimentary- non-functional', 6 Per.7_length / length_not_reduced_compared_to_pereonite_6 'length reduced- less than pereonite 6' pereonite_7_completely_undeveloped, 7 Head_anterolat._projections / antennae_emerging_directly_from_head squat_anterolat._projections_supporting_antennae robust_rounded_anterolat._projections_supporting_antennae 'long- thin anterolat. projections supporting antennae', 8 Head_lobe_on_ventrolateral_margin / absent_in_lateral_view present_in_lateral_view, 9 Spination_pattern_on_pers.1_to_4 / identical_for_males_and_females sexually_dimorphic, 10 Per.1_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 11 Per.1_female_anterolat._spines_length / 'elongate- length near width of pereonite 1' 'short- length near pereonite 1 length' 'tiny- length distinctly less than pereonite 1 length', 12 Per.1_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 13 Per.2_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 14 Per.2_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 15 Per.3_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 16 Per.3_female_paired_lateral_spines / no_lateral_spines_or_tubercles paired_lateral_simple_spines paired_lateral_pedestal_spines paired_lateral_rook_spines paired_lateral_tubercles, 17 Per.4_female_paired_dorsal_spines_or_tubercles / no_paired_dorsal_spines_or_tubercles paired_dorsal_simple_spines paired_dorsal_pedestal_spines paired_dorsal_rook_spines paired_dorsal_tubercles, 18 Per.4_female_paired_anterolat._spines_or_tubercles / no_anterolat._spines_or_tubercles anterolat._simple_spines anterolat._pedestal_spines anterolat._rook_spines anterolat._tubercles, 19 Per.4_female_with_posterolat._spines_or_tubercles / posterolat._spines_or_tubercles_absent posterodorsal_spines_or_tubercles, 20 Per.4_female_anterolat._spines_length / 'elongate- length near width of pereonite 1' 'short- length near pereonite 1 length' 'tiny- length distinctly less than pereonite 1 length', 21 Per.5_anterodorsal_spines_or_tubercles / anterodorsal_spines_or_tubercles_absent anterodorsal_rook_spines anterodorsal_pedestal_spines anterodorsal_simple_spines anterodorsal_tubercles, 22 Per.5_posterodorsal_spines_or_tubercles / no_posterodorsal_spines_or_tubercles posterodorsal_simple_spines posterodorsal_rook_spines posterodorsal_pedestal_spines posterodorsal_tubercles, 23 Per.5_spines_or_tubercles_lateral_row / no_lateral_row_spines_or_tubercles lateral_row_spines lateral_row_tubercles, 24 Per.6_dorsal_spines_or_tubercles / no_dorsal_spines_or_tubercles dorsal_simple_spines dorsal_rook_spines dorsal_pedestal_spines dorsal_tubercles, 25 PL_1_lateral_spines_or_tubercles / no_lateral_spines_or_tubercles lateral_simple_spines lateral_pedestal_spines lateral_rook_spines lateral_tubercles, 26 PLtn._dorsal_surface_axial_ridge / 'smooth arched- not separated from lateral fields' 'weakly vaulted- separated from lateral fields by shallow elongate concavities' 'strongly vaulted- separated from lateral fields by distinct indented lines', 27 PLtn._axial_ridge_length / axial_ridge_not_truncated axial_ridge_truncated, 28 PLtn._axial_ridge_distal_tip / medially_smoothly_arched 'medially incised- bilobed posteriorly' 'medially indented- hollow posteriorly', 29 PLtn._at_uropod_insertion / produced_from_margin indented_into_margin inflected, 30 PLtn._with_dorsal_spines_or_tubercles / dorsal_spines_or_tubercles_absent dorsal_tubercles dorsal_rook_spines dorsal_simple_spines dorsal_pedestal_spines, 31 PLtn._spines_or_tubercles_posterolat._marg. / spines_or_tubercles_absent spines_or_tubercles_present, 32 PLtn._type_of_spines_posterolat._marg. / simple_spines pedestal_spines rook_spines tubercles, 33 PLtn._terminal_margin_with_spines_or_tubercles / spines_or_tubercles_absent tubercles paired_simple_spines paired_pedestal_spines, 34 Antennula_num.__articles / two_or_less_articles 3_articles 4_articles 5_articles 6_or_more_articles, 35 Antennula_article_2_length / subequal_to_article_1_length less_than_2_times_article_1_length approximately_2_times_article_1_length greater_than_three_times_article_1_length, 36 Antennula_article_2_insertion / inserting_on_article_1_anteriorly anterodorsally dorsally, 37 Antennula_article_2_shape / 'straight- not curved at proximal insertion' strongly_curved_at_proximal_insertion, 38 Antennula_article_2_ventromedial_setae / 2_or_less_ventromedial_setae 3_ventromedial_setae 4_ventromedial_setae 5_ventromedial_setae 6_ventromedial_setae, 39 Antennula_article_2_distal_tip_shape / 'produced distally beyond insertion of next article- distal article inserting subapically' 'not produced distally- next article inserting apically', 40 Antennula_article_2_lateral_margin / robust_setae_absent row_robust_setae one_robust_seta, 41 Antennula_distal_articles_length / 'elongate- longer than article 2' subequal_to_article_2 'small- shorter than article 2' 'tiny- less than 0.3 article 2 length', 42 Antennula_article_3_length / article_3_longer_than_article_4 similar_length_to_article_4 'terminal- no distal articles', 43 Antennula_article_3_shape / 'elongate and tubular- much longer than wide' 'short and tubular- length near width' 'squat- wider than long', 44 Antennula_terminal_article_length / shorter_than_penultimate_article longer_than_penultimate_article, 45 Antenna_article_3_cuticle_smooth / smooth granulated, 46 Antenna_female_article_3_ventromedial_spines / no_ventromedial_spines ventromedial_spines, 47 Labrum_knobs / absent indistinct_low_projection present, 48 Maxilliped_palp_articles_width / article_2_wider_than_article_3 'articles 2-3 width subequal' article_3_wider_than_article_2, 49 Maxilliped_epipod_spines / epipod_spines_present epipod_spines_absent, 50 Per.bases_proximal_part_of_shaft / projecting_dorsally_above_shoulder dorsal_projection_absent, 51 Per.bases_proximal_shoulder / spines simple_spines bifid_spines rook_spines, 52 Per.I_shape / weakly_carposubchelate strongly_carposubchelate, 53 Per.I_merus_dorsal_surface / without_robust_setae with_distinctly_robust_setae, 54 Per.I_carpus_palm_vent._marg. / palm_or_distinctly_wider_part_absent proximally_expanded medially_expanded medially_expanded, 55 Per.I_carpus_vent._marg._robust_elongate_setae / 1_elongate_robust_seta 2_elongate_robust_setae row_similar_robust_setae 'fine setae only (no robust seta)', 56 Per.I_carpus_vent._marg._short_proximal_setae / several_setae_proximal_to_elongate_setae one_seta_proximal_to_elongate_seta setae_proximal_to_elongate_setae_absent, 57 Per.I_carpus_vent._marg._distal_setae_on_palm / several_distal_setae_on_palm 1_distal_seta_on_palm distal_setae_on_palm_absent, 58 Per.I_propodus_vent._marg._setae / several_robust_setae robust_setae_absent one_robust_seta, 59 Per.VII_presence / 'present- shorter than Per.VI' absent, 60 Pereopods_II, 61 PL.II_operculum_shape / broadly_triangular 'flap-like- much longer than wide' 'narrow proximal neck- almost circular posteriorly' 'narrow proximal neck- laterally convex- broadening posteriorly to rounded angles- posterior margin weakly convex' 'narrow proximal neck- laterally convex- broadening posteriorly to rounded angles- posterior margin weakly concave', 62 PL.II_operculum_setae / plumose_setae_absent plumose_setae_present, 63 Uropods_num.__articles / one_article 'two articles- protopod distinct', 64 Female_uropods_length_relative_to_pleotelson / extending_near_posterior_margin_of_pleotelson 'minute- not extending near posterior margin of pleotelson' 'elongate- extending beyond posterior margin of pleotelson', 65 Uropods_female_length_or_width / less_than_five_times_as_long_as_wide greater_than_or_equal_to_five_times_as_long_as_wide, 66 Uropods_female_length_or_length_of_pleotelson / less_than_0.4_length_of_pleotelson greater_than_or_equal_to_0.4_length_of_pleotelson, 67 Per.1_male_anterolat._spines_length / 'extremely elongate- length distinctly greater than width of pereonites' 'elongate- length near width of pereonite 1' 'short- length near length of pereonite 1' 'tiny- length distinctly less than length of pereonite 1', 68 Per.2_male_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 69 Per.3_male_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 70 Per.4_male_with_paired_anterolat._spines / no_paired_anterolat._spines 1_pair_of_anterolat._simple_spines 1_pair_of_anterolat._pedestal_spines 1_pair_of_anterolat._rook_spines 1_pair_of_anterolat._tubercles, 71 Per.4_male_length_of_anterolat._spines / 'elongate- length near width of pereonite 1' 'short- length near length of pereonite 1' 'tiny- length distinctly less than length of pereonite 1', 72 Per.4_male_posterolat._spines_or_tubercles / no_posterolat._spines_or_tubercles posterolat._spines_or_tubercles, 73 Antenna_flagellum_form_male / 'decreasing in width distally- tubular- longer than wide' 'some segments expanded- wider than long- with more setae on distal margins than juveniles or females', 74 PL.I_male_setae_on_distal_margin / setae_absent simple_setae blunt_setae plumose_setae, 75 PL.II_male_position_of_setae_on_protopod / setae_on_lateral_or_distal_margin setae_on_distal_margin_only setae_absent, 76 PL.II_stylet_length / 'short- reaching end of protopod' 'elongate- reaching beyond end of protopod', 77 PL.II_stylet_tip_shape / 'thread-like' thick_and_blunt thin_with_medial_blade, 78 Uropods_male_length_relative_to_pleotelson / not_extending_near_posterior_margin_of_pleotelson 'extending near posterior margin of pleotelson (+ or - 10pct)' extending_beyond_posterior_margin_of_pleotelson, 79 Uropods_male_length_or_pleotelson_length / less_than_0.5_length_of_pleotelson greater_than_0.5_length_of_pleotelson, ; MATRIX [ 10 20 30 40 50 60 70 80 ] [ . . . . . . . . ] Contrarimesus_curtispinis 22222111?121121511-111111211111-154311213212?1?322?2211331?1322312????????????? Contrarimesus_franklinae 22222211?2111111123111111211111-15431321311121?22212211321114223??222231?2111?? Cornuamesus_biscayensis 2222124??-----------11111211112115431-2131112-?11??21123111?--1---222231?213331 Cornuamesus_longiramus 222211412121111111-114111211112115431321311211?11122111213115113222222211212332 Fortimesus_brevispinis 2222111??5325252523??1111211221-354312223112???22???????????221211------------- Fortimesus_concinnus 2222111??-----------55253211241-354312212112?-?32??2112311??--1---252212?211111 Fortimesus_consanguineus 22221111?132121211-111111211211-154312223111???32??2212311??421111------------- Fortimesus_cornutus 2222121??-----------11111211221-3543152231121-?12??221231121--1---2221-1?211111 Fortimesus_gigas 2222121?1131111111-111111211211-35431-223111?1?32??2112311??3112114111-11211111 Fortimesus_profundicolus 2222121??{25}22{25}25221-251111211221-344314223112?1?32??2212311??411111------------- Fortimesus_robustus 2222111??135155151-251111211211-153312213111?1?32??2212311??421211??????121111? Fortimesus_scabriusculus 2222111??1-1111111-211211211211-354312223212?1?22??2?12311?1221211------------- Fortimesus_thomsoni 2222111??-----------15253211211-3?????2??????-??221211131?12--1----11232?1???11 Fortimesus_zuluensis 22221211?521555551-151111211211-1?431??1311?21?22??211231111??1111------------- Gracilimesus_angustus 22221311?-----------111112112121144?112132121-?32212?1122331--1---3111-1?1???21 Gracilimesus_celticensis 222213112121111111-111111211212214431121321211?12212?1122331411111211221?111121 Gracilimesus_corniculatus 22221111?-----------1111121121221543112132111-?1221211132331--1---2111-12212121 Gracilimesus_gorbunovi 2222111??121111111-111111111212115431?21221111??2?????????????11?1------------- Gracilimesus_gorbunovi_sensu_Svavarsson 2222121111211111122111111111212115431121311111?22212111323114111111112111211121 Gracilimesus_hanseni 2222131111211111122111111211212215431121321111?12212211123314211112112112211121 Gracilimesus_insignis 22221211?-----------1111121121211543122131111-?12212111323?1--1---11121112???31 Gracilimesus_orientalis 2222111??-----------1111121121211542112131111-?12??21112232?--1---211211?211121 Gracilimesus_tenuispinis 22221111?1211111121111111111212115431121311111?122?2111323?1421311------------- Gracilimesus_tropicalis 222213112131111111-11111121121211?431121?????1?12212?112133?411111411231?2???21 Haplomesus_bifurcatus 22221211?-----------11111313211-1?431221??1?1-?32??2?1122112--1---2121-11213111 Haplomesus_ornatus 22221211?-----------11111322211-1543122131112-?3221211122111--1---2121-112???11 Haplomesus_quadrispinosus 22221211?121111111-111111311211-15431321311121?32212111221113?1111------------- Haplomesus_quadrispinosus_sensu_Brandt 22221111?-----------11111322211-1543122131112-?2221211122321--1---1121-11213111 Heteromesus_drachi 1222-112?-----------111??3111?1-?34323214121?-122123?31211?1--1---4221-11112331 Heteromesus_frigidus 1222-11?1131111111-111111311111-12432211433-21322113?31211133213?{12}4112?21112332 Heteromesus_granulatus 1222-11??531111111-111111311111-12432311433-21322123?41211133213114??????????21 Heteromesus_schmidtii 1222-11?1532555151-111111311111-12432211433-22322123?41221123213?2------------- Heteromesus_similis 1222-112?????????????????3?11????4????2?????2?122123?31211?3??1???4221-211???31 Heteromesus_spinosus 1222-11??4234343123211244311152244432211421122322143?41211123213?2------------- Ischnomesus_antarcticus 1111-1111-----------111??1112?1-?542112111121-?1221312211111--2---3112221422231 Ischnomesus_bispinosus 1111-1111131111111-111111111211-15421221211111112?13221112112?23113111-11223131 Stylomesus_hexapodus 1122-231?131111111-111111111211-113313?14???21?22213?2121133412322------------- Stylomesus_menziesi 1122-13?1-----------11111111211-1533122132112-??2??3?11213?1--2----111-1?213122 Stylomesus_sarsi 1122-231????????????11111211211-1?321221????2??12??31112111242232?4211-12211132 ; END; BEGIN TREES; TITLE Tb8706; LINK TAXA = Taxa1; TRANSLATE 1 Stylomesus_sarsi, 2 Stylomesus_menziesi, 3 Stylomesus_hexapodus, 4 Contrarimesus_curtispinis, 5 Ischnomesus_bispinosus, 6 Ischnomesus_antarcticus, 7 Contrarimesus_franklinae, 8 Fortimesus_zuluensis, 9 Fortimesus_thomsoni, 10 Gracilimesus_tenuispinis, 11 Fortimesus_scabriusculus, 12 Fortimesus_robustus, 13 Haplomesus_quadrispinosus_sensu_Brandt, 14 Haplomesus_quadrispinosus, 15 Haplomesus_bifurcatus, 16 Fortimesus_profundicolus, 17 Haplomesus_ornatus, 18 Gracilimesus_orientalis, 19 Gracilimesus_insignis, 20 Gracilimesus_gorbunovi_sensu_Svavarsson, 21 Gracilimesus_gorbunovi, 22 Fortimesus_gigas, 23 Fortimesus_consanguineus, 24 Fortimesus_cornutus, 25 Fortimesus_concinnus, 26 Fortimesus_brevispinis, 27 Cornuamesus_longiramus, 28 Cornuamesus_biscayensis, 29 Gracilimesus_tropicalis, 30 Gracilimesus_corniculatus, 31 Gracilimesus_angustus, 32 Gracilimesus_celticensis, 33 Gracilimesus_hanseni, 34 Heteromesus_spinosus, 35 Heteromesus_similis, 36 Heteromesus_schmidtii, 37 Heteromesus_granulatus, 38 Heteromesus_frigidus, 39 Heteromesus_drachi; TREE Fig._3B = [&R] ((6,5)Ischnomesus,((2,(3,1))Stylomesus,(((12,8),((23,(26,(24,16))),(25,9),22,11))Fortimesus,(((33,30),(29,(32,31)),21,20,19,18,10)Gracilimesus,((17,15,14,13)Haplomesus,((7,4)Contrarimesus,((28,27)Cornuamesus,(39,(35,(38,37,36,34)))Heteromesus))))))); END; BEGIN TREES; TITLE Tb8705; LINK TAXA = Taxa1; TRANSLATE 1 Stylomesus_sarsi, 2 Stylomesus_menziesi, 3 Stylomesus_hexapodus, 4 Contrarimesus_curtispinis, 5 Ischnomesus_bispinosus, 6 Ischnomesus_antarcticus, 7 Contrarimesus_franklinae, 8 Fortimesus_zuluensis, 9 Fortimesus_thomsoni, 10 Gracilimesus_tenuispinis, 11 Fortimesus_scabriusculus, 12 Fortimesus_robustus, 13 Haplomesus_quadrispinosus_sensu_Brandt, 14 Haplomesus_quadrispinosus, 15 Haplomesus_bifurcatus, 16 Fortimesus_profundicolus, 17 Haplomesus_ornatus, 18 Gracilimesus_orientalis, 19 Gracilimesus_insignis, 20 Gracilimesus_gorbunovi_sensu_Svavarsson, 21 Gracilimesus_gorbunovi, 22 Fortimesus_gigas, 23 Fortimesus_consanguineus, 24 Fortimesus_cornutus, 25 Fortimesus_concinnus, 26 Fortimesus_brevispinis, 27 Cornuamesus_longiramus, 28 Cornuamesus_biscayensis, 29 Gracilimesus_tropicalis, 30 Gracilimesus_corniculatus, 31 Gracilimesus_angustus, 32 Gracilimesus_celticensis, 33 Gracilimesus_hanseni, 34 Heteromesus_spinosus, 35 Heteromesus_similis, 36 Heteromesus_schmidtii, 37 Heteromesus_granulatus, 38 Heteromesus_frigidus, 39 Heteromesus_drachi; TREE Fig._3A = [&R] ((6,5),((2,(3,1)),(((12,8),((23,(26,(24,16))),(22,(11,(25,9))))),(((21,(29,(32,31))),(18,((33,30),20,19,10))),((17,15,14,13),((7,4),((28,27),(39,(35,(38,37,36,34)))))))))); END;