#NEXUS

[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on April 24, 2024; 18:55 GMT

TreeBASE (cc) 1994-2008

Study reference:
Jamieson B. 1988. On the phylogeny and higher classification of the Oligochaeta.
Cladistics, 4: 367-410.

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S363]

BEGIN TAXA;
        TITLE  Taxa1;
        DIMENSIONS NTAX=50;
        TAXLABELS
            Ailoscolex_lacteospumosus
            Alluroides_pordagei
            Alma_nilotica
            Biwadrilus_bathybates
            Brinkhurstia_americanus
            Callidrilus_scrobifer
            Celeriella_bursata
            Criodrilus_lacuum
            Cryptodrilus_resticus
            Desmogaster_doriae
            Dichogaster_bolaui
            Dichogaster_oraedividitis
            Digaster_bradburyi
            Diplotrema_fragilis
            Diporochaeta_intermedia
            Diporodrilus_pilosus
            Drilocrius_hummelincki
            Eiseniella_teraedra
            Enchytraeidae
            Eudrilus_eugeniae
            Eukerria_garmani_argentinae
            Glossoscolex_paulistus
            Glyphidrilocrius_ehrhardti
            Glyphidrilus_weberi
            Haplotaxis_violaceus
            Hemigastrodrilus_monicae
            Heteroporodrilus_tryoni
            Hormogaster_redii
            Komarekiona_eatoni
            Kynotus_darwini
            Lampito_sylvicola
            Lumbriculidae
            Lumbricus_terrestris
            Lutodrilus_multivesiculatus
            Megascolides_australis
            Microchaetus_benhami
            Microscolex_georgianus
            Moniligaster_troyi
            Nemertodrilus_griseus
            Ocnerodrilus_occidentalis
            Octochaetus_multiporus
            Perionyx_excavatus
            Pheretima_montana
            Pontoscolex_corethrurus
            Sparganophilus_tamesis
            Standeria_tranvaalensis
            Syngenodrilus_lamuensis
            Tiguassu_reginae
            Tubificidae
            Vignysa_popi
            ;
END;

BEGIN CHARACTERS;
[! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2023]

	TITLE Table_2;
	LINK TAXA = Taxa1;
	DIMENSIONS NCHAR=68;
	FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4 5 6 7 8 9 A B C D E" MISSING=? GAP= -;
	CHARSTATELABELS
		1  Prostomium / 'attached (P)' 'separate (A)',
		2  Lateral_lines / 'present (P)' 'absent (A)',
		3  Dorsal_pores / 'absent (P)' 'present (A)',
		4  Caudal_TS / 'not quadrangular (P)' 'quadrangular (A)',
		5  Anus / 'terminal (P)' 'dorsal (A)',
		6  Copulatory_alae / 'absent (P)' 'present (A)',
		7  Claspers_with_terminal_male_por / 'absent (P)' 'present (A)',
		8  Genital_lobes_or_claspers / 'without lateral male pores (P)' 'with lateral male pores (A)',
		9  Longitudinal_tubercula_pubertat / 'absent (P)' 'present (A)',
		10  Somatic_setae / '4 bundles (P)' 'perichaetine (A)',
		11  Penial_setae / 'absent (P)' 'present (A)',
		12  'Long-grooved genital setae' / 'absent (P)' 'present (A)',
		13  'Prostate-like setal glands' / 'absent (P)' 'present (A)',
		14  Clitellum / 'unilayered (P)' 'multilayered (A)',
		15  Dorsal_vessel / 'extending onto pharynx (P)' 'or not (A)',
		16  Subneural_vessel / 'absent (P)' 'present (A)',
		17  Intestine_with_anterior_thicken / 'absent (P)' 'present (A)',
		18  Intestinal_caeca / 'absent (P)' 'present (A)',
		19  Intestinal_typhlosole / 'absent (P)' 'present (A)',
		20  Enterosegmental_organs / 'absent (P)' 'paired (A)',
		21  Nephridia / 'holonephric (P)' 'meronephric (A)',
		22  Nephridial_bladder / 'absent (P)' 'present (A)',
		23  Nephridial_caeca / 'absent (P)' 'present (A)',
		24  Caudal_nephridia / 'exonephric (P)' 'enteronephric (A)',
		25  Testes / in_segment_10_or_homeotic_equiv 'not in 10 (A)',
		26  Testes / in_segment_11_or_homeotic_equiv 'not in 11 (A)',
		27  Male_pores / '2 per side (P)' '1 by fusion (A)',
		28  Muscular_male_bursa / 'absent (P)' 'present (A)',
		29  Glandular_male_bursa / 'absent (P)' 'present (A)',
		30  Ovaries / 'in segment 12 (P)' 'not in 12 (A)',
		31  Ovaries / 'in segment 13 (P)' 'not in 13 (A)',
		32  Ovaries / 'multistringed (P)' 'single-stringed (A)',
		33  Ovaries / 'free (P)' 'in septal chambers (A)',
		34  Spermathecae / 'present (P)' 'absent (A)',
		35  Spermathecae / 'not multiple (P)' 'multiple (A)',
		36  Spermathecae / 'without oviducal link (P)' 'oviducal link present (A)',
		37  'Supra-oesophageal vessel' / 'absent (P)' 'on oesophagus only (Al)' 'oesophago-pharyngeal (A2)',
		38  Commissural_blood_vessels / 'throughout (P)' 'discrete hearts in 10 (Al)' '11 (A2)' '12 (A3)' '13 (A4)',
		39  Segments_of_oesophageal_gizzard / '0 (P)' '1 (Al)' '2 (A2)' '3 (A3)' '4 (A4)' '12 (A5)',
		40  1st_oesophageal_gizzard_in_segm / 'absent (P)' '5 (Al)' '6 (A2)' '7 (A3)' '8 (A4)' '13 (A5)' '14 (A6)',
		41  1st_intestinal_segment / '9 (P)' '10 (Al)' '11 (A2)' '12 (A3)' '13 (A4)' '14 (A5)' '15 (A6)' '16 (A7)' '17 (A8)' '21 (A9)' '23 (A10)',
		42  'Last male pore(s)' / 'posterior to last testis-segmen' '2 (Al)' '3 (A2)' '4 (A3)' '5 (A4)' '6 (A5)' '7 (A6)' '8 (A7) 9 (A8)' '11 (A9)' '12 (A10)' '13 (All) segments',
		43  Setal_interval / 'cd=ab (P)' 'cd>ab (A)',
		44  Setal_interval / 'cd=ab (P)' 'cd<ab (A)',
		45  Caudal_nephrostomes / '1 per side (P)' 'many per side (A)',
		46  Caudal_nephridia / 'present (P)' 'none (A)',
		47  'Testis-sacs' / 'absent (P)' 'segmental (A)',
		48  'Testis-sacs' / 'absent (P)',
		49  'Testis-sacs' / 'absent (P)' 'capping the testes (A)',
		50  Seminal_vesicles / 'extensive (P)' 'short (A)',
		51  Seminal_vesicles / 'present (P)' 'absent (A)',
		52  Male_pores / 'anterior on clitellum (P)' 'anterior to clitellum (A)',
		53  Male_pores / 'anterior on clitellum (P)' 'posterior to clitellum (A)',
		54  Tubular_prostates / 'absent (P)' 'present (A)',
		55  'Tubulo-racemose or racemose pro' / 'absent (P)' 'present (A)',
		56  Capsular_prostates / 'absent (P)' 'present (A)',
		57  Euprostates / 'absent (P)' 'present (A)',
		58  Spermathecae / 'adiverticulate (P)' 'seminal diverticulum (A)',
		59  Spermathecae / 'adiverticulate (P)' 'with non-seminal diverticulum (',
		60  Acanthrodrilin_prostate_arrange / 'absent (P)' 'present (A)',
		61  Microscolecin_prostate_arrangem / 'absent (P)' 'present (A)',
		62  Megascolecin_prostate_arrangeme / 'absent (P)' 'present (A)',
		63  Last_spermathecae / 'pretesticular (P)' 'testicular (A)',
		64  Last_spermathecae / 'pretesticular (P)' 'post-testicular (A)',
		65  Number_of_clitellar_segments / '3 (P)' '3.5-5 (Al )' '6 17 (A2)' '20 60 (A3)',
		66  Eudriloid_spermatheca / 'absent (P)' 'present (A)',
		67  Secondary_tube_of_sperm_acrosom / 'free (P)' 'with weak connectives (Al)' 'full connectives (A2)',
		68  Length_of_acrosome / 'very short (P)' 'short (Al)' 'moderate (A2)' 'long (A3)',
		;

MATRIX
[                                       10        20        30        40        50        60        ]
[                                        .         .         .         .         .         .        ]
Ailoscolex_lacteospumosus       01000000100101000010011000100101000002426A010000010010000000001030??
Alluroides_pordagei             000000000000000000000000011001000000030041000000010000010000000010??
Alma_nilotica                   0101101000100110101000000010010000102300C7000000010100000000000130??
Biwadrilus_bathybates           000010000010110000000000001001000100120021010000010100000000000030??
Brinkhurstia_americanus         000000000010000000000000011001000000030071000000001001000000000010??
Callidrilus_scrobifer           010110000000011100100000001001000010232175000000010000100000000130??
Celeriella_bursata              011000000100010000001000001001000000131286001000010011000100010010??
Criodrilus_lacuum               010110000001010110100000001011010100022143000000010000000000000030??
Cryptodrilus_resticus           011000000010010000001100001001000000131196100100010010100100010010??
Desmogaster_doriae              11000000000000010001011000000100100002CAE0000001001000010010000020??
Dichogaster_bolaui              011000000010010000101000001001000000142296000000010011000101000020??
Dichogaster_oraedividitis       0110000000000100001010000010010000001322A6000010010011000101000020??
Digaster_bradburyi              01100000000001000000100000100100000013220610000001001010010001001023
Diplotrema_fragilis             011000000010010000000000001001000000141166100000010011000101000010??
Diporochaeta_intermedia         010000000100010000000100001001000000131166000000010011000100010000??
Diporodrilus_pilosus            010000001000010010100100001001010000020073000000010100000000001020??
Drilocrius_hummelincki          010110010000011110100000001001000010222183000000010000000000000130??
Eiseniella_teraedra             01110000100001011010010000100101000002006100000001010000000000102023
Enchytraeidae                   00000000000000000000000010000010000000000000000000000000000000000000
Eudrilus_eugeniae               010000000000011100000000001001000001121165000010010010001100100121??
Eukerria_garmani_argentinae     010000000000011000000000011001000000120036000000010011000001000020??
Glossoscolex_paulistus          010000001000011000100110101101000000121258010010010000000000000020??
Glyphidrilocrius_ehrhardti      0101100100000110101000000010010000102400E5000000010000000000000130??
Glyphidrilus_weberi             01011100000001010010000000100100001012236D100000010010000000000130??
Haplotaxis_violaceus            00000000000000000000000000000010000000000000000001000000000000000020
Hemigastrodrilus_monicae        010000001000010000100000001001000000023253010000010000000000000130??
Heteroporodrilus_tryoni         0110000000100100000001000010010000001411661000000100101001000100102?
Hormogaster_redii               01000000100101010010011000100101000012327401000001000001000000013023
Komarekiona_eatoni              01000000100011000000010000100101000002124A000000010010000000000020??
Kynotus_darwini                 0100000000001101000001100011010000101211C4000010010100000000C00130??
Lampito_sylvicola               011000000100010000101001101001000000141176000000010010100100010020??
Lumbriculidae                   00000000000000?000000000000000??0000?000?0??000000000000000000??0011
Lumbricus_terrestris            01100000100101011010010000100101000002006301001001010000000000102023
Lutodrilus_multivesiculatus     01011100000011110010000000100101001002002B000000010000000000000130??
Megascolides_australis          0110000000000100000010010010010000000411A6100000010011000100010000??
Microchaetus_benhami            010000001010110000100110001001000000021359010000010010000000000130??
Microscolex_georgianus          010000000010010000000100001001000000131176100000010011000101000010??
Moniligaster_troyi              110000000000000100010110100001001000023980000001001000010010000010??
Nemertodrilus_griseus           010000000000010100000000001001000001121176000000010010001000000121??
Ocnerodrilus_occidentalis       010000000000010000000000001001000100120035000000101011000000100020??
Octochaetus_multiporus          0110000000000100001010010010010000001412A6001000010011000101000020??
Perionyx_excavatus              011000000110010100000000001001000000231266000000010010100100010010??
Pheretima_montana               01100000010001010110100100100100000014146600101001001010010001000023
Pontoscolex_corethrurus         010000001000010100100110101001000000131289100010000010000000000020??
Sparganophilus_tamesis          01001000100011000000000000100101000002000700000001000000000000002022
Standeria_tranvaalensis         000000000000000000000000011101000000030061000000001000000000000010??
Syngenodrilus_lamuensis         0?00000010000000000001000010010000000324310000100000010000000000?0??
Tiguassu_reginae                01000000100000???0000000100000100000?13230010000000100000000000020??
Tubificidae                     00000000000000000000000010011010000000000000000000000000100000100000
Vignysa_popi                    010000001000010010100000001001000000022263010000010000000000001020??
;
END;

BEGIN TREES;
      TITLE Tb5156;
      LINK TAXA = Taxa1;
         TRANSLATE
            1    Pheretima_montana,
            2    Lampito_sylvicola,
            3    Dichogaster_oraedividitis,
            4    Dichogaster_bolaui,
            5    Octochaetus_multiporus,
            6    Megascolides_australis,
            7    Diporochaeta_intermedia,
            8    Celeriella_bursata,
            9    Perionyx_excavatus,
            10   Digaster_bradburyi,
            11   Cryptodrilus_resticus,
            12   Heteroporodrilus_tryoni,
            13   Microscolex_georgianus,
            14   Diplotrema_fragilis,
            15   Eiseniella_teraedra,
            16   Lumbricus_terrestris,
            17   Komarekiona_eatoni,
            18   Ailoscolex_lacteospumosus,
            19   Diporodrilus_pilosus,
            20   Hemigastrodrilus_monicae,
            21   Vignysa_popi,
            22   Hormogaster_redii,
            23   Kynotus_darwini,
            24   Pontoscolex_corethrurus,
            25   Glossoscolex_paulistus,
            26   Microchaetus_benhami,
            27   Nemertodrilus_griseus,
            28   Eudrilus_eugeniae,
            29   Eukerria_garmani_argentinae,
            30   Ocnerodrilus_occidentalis,
            31   Criodrilus_lacuum,
            32   Lutodrilus_multivesiculatus,
            33   Callidrilus_scrobifer,
            34   Glyphidrilocrius_ehrhardti,
            35   Drilocrius_hummelincki,
            36   Glyphidrilus_weberi,
            37   Alma_nilotica,
            38   Sparganophilus_tamesis,
            39   Biwadrilus_bathybates,
            40   Syngenodrilus_lamuensis,
            41   Standeria_tranvaalensis,
            42   Brinkhurstia_americanus,
            43   Alluroides_pordagei,
            44   Desmogaster_doriae,
            45   Moniligaster_troyi,
            46   Tiguassu_reginae,
            47   Haplotaxis_violaceus,
            48   Lumbriculidae,
            49   Tubificidae,
            50   Enchytraeidae;
      TREE Fig._1B = [&R] ((50,49)Tubificata,(47,(48,(46,((45,44)Moniligastridae,(((43,(42,41))Alluroididae,40)Alluroidina,(((39,((((((37,36),33),35),31),34)Almidae,32)),38)Aquamegradrili,((30,29)Ocnerodrilidae,((28,27)Eudrilidae,(((((26,(22,20)),(25,24)Glossoscolecidae),((21,(19,(16,15))Lumbricidae),18)),17),((14,13),(12,(11,(10,(9,(8,(7,((6,(5,(4,3))),(2,1)))))))))Megascolecidae)))Terrimegadrili)Crassiclitellata)Opisthopora)Metagynophora))))Oligochaeta;
      TREE Fig._1A = [&R] ((50,49),(48,(47,(46,((45,44),(((43,(42,41)),40),(((39,((((((37,36),33),35),31),34),32)),38),((30,29),((28,27),(((((26,(22,20)),(25,24)),((21,(19,(16,15))),18)),17),((14,13),(12,(11,(10,(9,(8,(7,((6,(5,(4,3))),(2,1)))))))))))))))))Diplotesticulata));



END;