#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on April 25, 2024; 0:21 GMT TreeBASE (cc) 1994-2008 Study reference: Prendini L. 2000. Phylogeny and classification of the superfamily Scorpionoidea Latreille 1802 (Chelicerata, Scorpiones): an exemplar approach. Cladistics, 16: 1-78. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S652] BEGIN TAXA; TITLE Taxa1; DIMENSIONS NTAX=71; TAXLABELS Bioculus_caboensis Bioculus_comondae Bothriurus_bonariensis Bothriurus_coriaceus Brachistosternus_alienus Brachistosternus_ehrenbergii Brachistosternus_ferrugineus Cazierius_gundlachii Cazierius_scaber Centromachetes_obscurus Centromachetes_pocockii Centruroides_gracilis Cercaphonius_queenslandae Cercophonius_squama Chaeritus_granosus Cheloctonus_crassimanus Cheloctonus_jonesii Chiromachetes_fergusoni Chiromachetes_tirupati Chiromachus_ochropus Didymocentrus_hasethi Didymocentrus_lesueurn Diplocentrus_gertschi Diplocentrus_mexicanus Habibiella_gaillardi Hadogenes_tityrus Hadogenes_troglodytes Hemiscorpius_lepturus Hemiscorpius_maindroni Heterometrus_spinifer Heterometrus_swammerdami Heteronebo_granti Heteronebo_jamaicae Heteroscorpion_goodmani Heteroscorpion_opisthacanthoides Iomachus_laeviceps Iomachus_politus Liocheles_australasiae Liocheles_waigiensis Lisposoma_elegans Lisposoma_josehermana Nebo_hierichonticus Nebo_poggesii Oiclus_purvesii Opisthacanthus_elatus Opisthacanthus_madagascariensis Opisthacanthus_validus Opistophfhalmus_boehmi Opistophthalmus_capensis Opistophthalmus_holmi Orobothriurus_alticola Orobothriurus_crassimanus Palaeocheloctonus_pauliani Pandinus_cavimanus Pandinus_dictator Pandinus_imperator Phoniocercus_piclus Phoniocercus_sanmartini Scorpio_maurus_kruglovi Scorpio_maurus_mogadorensis Tarsoporosus_flavus Tehuankea_moyanoi Thestylus_glasioui Timogenes_dorbignyi Timogenes_mapuche Torsoporosus_kugleri Urodacus_novaehollandiae Urodacus_yaschenkoi Urophonius_granulatus Urophonius_iheringii Vachonia_martinezi ; END; BEGIN TAXA; TITLE Taxa2; DIMENSIONS NTAX=37; TAXLABELS Bioculus Bothriurus Brachistosternus Buthidae Cazierius Centromachetes Cercophonius Chaerilus Cheloctonus Chiromachetes Chiromachus Didymocentrus Diplocentrus Habibiella Hadogenes Hemiscorpius Heterometrus Heteronebo Heteroscorpion Iomachus Liocheles Lisposoma Nebo Oiclus Opisthacanthus Opistophthalmus Orobothriurus Pandinus Phoniocercus Scorpio Tarsoporosus Tehuankea Thestylus Timogenes Urodacus Urophonius Vachonia ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2591] TITLE Table_3; LINK TAXA = Taxa1; DIMENSIONS NCHAR=115; FORMAT DATATYPE=Standard SYMBOLS= "0 1 2 3 4" MISSING=? GAP= -; CHARSTATELABELS 1 'Carapace, Lateral ocelli, number of pairs' / more_than_three three two, 2 Median_ocular_tubercle / raised shallow., 3 Median_notch / absent shallow strongly_excavated, 4 Median_longitudinal_furrow / 'broad, shallow, without suture' 'narrow, suturiform.', 5 Median_longitudinal_furrow / 'well-developed' obsolete., 6 'Antero-ocular depression' / absent present., 7 Posterior_sutures / absent present., 8 'Nongranular surfaces of prosoma, mesosoma, metasoma, and legs' / smooth distinctly_punctate., 9 'Sternum, Sternum shape' / subtriangular subpentagonal transverse equilateral_pentagonal., 10 'Chelicerae, Cheliceral movable finger, number of subdistal teeth' / one two., 11 'Cheliceral movable finger, distal external and distal internal teeth' / 'subequal, with distal external tooth only slightly smaller than' 'unequal, with distal external tooth considerably smaller than d', 12 'Cheliceral fingers, secondary serrations' / absent present., 13 'Cheliceral coxae, scaphotrix (stridulatory setae) on dorsointernal surfaces' / absent present., 14 'Cheliceral coxae, trichopae (chemoreceptive lamelliform setae) on internal surfa' / absent present., 15 'Pedipalp Ornamentation, Patella, dorsal surface' / 'flat, dorsomedian, and dorsoexternal carinae in same axis' 'convex, dorsomedian carina raised above horizontal axis of dors', 16 'Patella, dorsoexternal carina' / distinct obsolete., 17 'Patella, externomedian carina' / continuous_from_proximal_to_distal_edges 'discontinuous, interrupted two-thirds along.', 18 'Patella, anterior process' / absent present., 19 'Chela, number of carinae' / 8. 10., 20 'Chela (male), dorsal secondary carina' / distinct obsolete., 21 'Chela (female), dorsal secondary carina' / distinct obsolete., 22 'Chela, dorsal secondary carina' / 'extending full way across dorsal surface, subdigital carina ves' dorsal_secondary_carina_extending_part_way_across_dorsal_surfac, 23 'Chela, digital carina' / distinct obsolete., 24 'Chela, disproportionate development of dorsal secondary and external secondary c' / absent present., 25 'Chela, disproportionate development of external secondary carina only, relative' / absent present., 26 'Chela, ventroexternal carina' / distinct obsolete., 27 'Chela, ventroexternal carina' / 'parallel to longitudinal axis of chela, distal edge connected t' 'parallel to longitudinal axis of chela, distal edge disconnecte' 'oblique to longitudinal axis of chela, distal edge disconnected' 'oblique to longitudinal axis of chela, distal edge disconnected', 28 'Chela, ventromedian carina' / vestigial_or_obsolete distinct, 29 'Chela, ventrointernal carina' / 'more strongly developed than internomedian carina, which may be' equally_or_less_strongly_developed_than_internomedian_carina., 30 'Chela, ventrointernal carina' / 'equally or more strongly developed than internomedian carina, w' 'less strongly developed than internomedian carina, often obsole', 31 'Chela (male), secondary sexual structure' / absent 'hook-like apophysis' 'semicircular, rimmed depression', 32 'Chela (male), hook-like secondary sexual structure with granular ridge at base o' / present absent., 33 'Chela fingers, number of rows of primary denticles' / single 'double, often fused at the base' multiple, 34 'Chela fingers, dentate margin' / entire markedly_scalloped., 35 'Chela (male), lobe of movable finger' / 'absent or at most weakly developed, close to base movable finge' 'welldeveloped, almost midway along movable finger (if dentate m' 'well-developed, almost midway along movable finger, with a dist', 36 'Chela (male, also applicable in females with welldeveloped lobe on movable finge' / absent present., 37 'Chela (female), lobe of movable finger' / 'absent or at most weakly developed, close to base of movable fi' 'well-developed, almost midway along movable finger, with distin', 38 'Chela (female), sharp conical tooth at base of movable finger and corresponding' / absent present., 39 'Trichobothria, Femur, number e r' / one two four, 40 'Femur, position 7 i' / internal dorsal., 41 'Patella, position 7. d2' / dorsal internal., 42 Patella_v_7 / absent three_prolaterals three_or_more_retrolaterals, 43 'Patella, number v r' / absent single_row_of_3 'single row of 4-20' 'two or more rows, with more than 30.', 44 'Patella, position distal v 7' / ventral external., 45 'Patella, number e r' / 7. '13, rarely 12' 14_or_more, 46 'Chela, number i r' / one two three_or_more, 47 'Chela, position r- it' / basal_or_midfinger distal., 48 'Chela, position r- ib and it' / basal midfinger., 49 'Chela, number V r. 1' / 2. 4. 5. 6_or_more, 50 'Chela, distance between r- V2 and V3' / normal widely_separated., 51 'Chela, position r Db' / external_surface dorsal_surface., 52 'Chela, position 7-Dt' / 'manus, at proximal end' 'manus, mid-length or slightly less than mid-length' 'manus, distal half, near base of fixed finger' proximal_end_of_fixed_finger, 53 'Chela, external surface with accessory 7' / absent '10-15' more_than_20, 54 'Chela, position 7- Eb3' / 'in line with Ebl-Eb2 axis' 'distal to Ebl-Eb2 axis.', 55 'Chela, position 7 Est' / distal midpalm., 56 'Chela, position 7-Et2' / external_surface ventral_surface., 57 'Chela, number d 7-' / two four, 58 'Chela, position 7. db' / fixed_finger manus., 59 'Chela, position r db' / dorsal internal., 60 'Chela, position 7 dsb' / 'below db-dst axis' 'in line with db-dst axis.', 61 'Chela, position of 7- eb' / proximal_region_of_fixed_finger 'manus, behind point of articulation between fixed and movable f', 62 'Chela, position of 7. esb' / 'manus, behind point of articulation between fixed and movable f' 'midway along fixed finger, in line with eb-est-et axis.', 63 'Legs, Retrolateral pedal spurs' / present absent., 64 Prolateral_pedal_spurs / present absent., 65 'Telotarsi, laterodistal lobes' / 'truncated, base of median dorsal lobe flush' 'rounded, notches at base of median dorsal lobe' 'secondarily truncated, notches at base of median dorsal lobe', 66 'Telotarsi, laterally flattened' / absent present., 67 'Telotarsi 1, numerous pores ventrally and prolaterally' / absent present., 68 'Telotarsi, well-developed ventromedian row of setae' / spiniform setiform absent, 69 'Telotarsi 1-1V, ventrosubmedian setae distribution' / 'setiform on I-IV' 'setiform on 1 or 1-11, spiniform on 111-IV' 'spiniform (or secondarily setiform) on I-IV', 70 'Telotarsi, ventrosubmedian setae type' / stout_spiniform slender_spiniform few_secondarily_setiform numerous_secondarily_setiform, 71 'Telotarsi IV, prolateral row of ventrosubmedian spiniform setae' / present absent., 72 'Basitarsi 1-11, retrolateral row of macrochaete setae' / absent spiniform 'setiform, sand comb', 73 'Basitarsi 1-11, reduction in the number of macrochaete setae in the retrolateral' / absent present, 74 Stridulatory_surface_on_opposing_surfaces_of_coxae_of_pedipalps_and_first_walkin / absent partially_developed fully_developed, 75 Stridulatory_surface_on_opposing_surfaces_of_coxae_of_pedipalps_and_first_walkin / 'rasp on pedipalpal coxa, scraper on coxa of first walking leg' 'scraper on pedipalpal coxa, rasp on coxa of first walking leg.', 76 'Maxillary lobes, shape of first pair' / roundedtruncate_anteriorly tapering_anteriorly., 77 'Reproductive Anatomy, Embryonic development' / apoikogenic katoikogenic., 78 Ovariuterine_follicles / sessile stalked., 79 Testis / straight coiled., 80 'Genital opercula (female)' / separated loosely_joined fused, 81 'Genital opercula (male)' / separated loosely_joined., 82 'Hemispermatophore and Paraxial Organ, Hemispermatophore' / flagelliform fusiform lamelliform, 83 'Hernispermatophore, truncal flexure' / absent present., 84 'Paraxial organ, internobasal reflection of sperm duct' / absent present., 85 'Paraxial organ, internal wall of sperm duct' / simple with_semilunar_shelf., 86 'Hemispermatophore, distal lamina' / smooth with_prominent_crest., 87 'Hemispermatophore, distal lamina' / simple with_complex_differentiation., 88 'Hemispermatophore, lamellar hook and median lobe' / separate fused., 89 'Hemispermatophore, position of lamellar hook' / basal distal, 90 'Hemispermatophore, lamellar hook' / single double, 91 'Hemispermatophore, spines in capsular region' / absent present., 92 'Hemispermatophore, sclerotised mating plug' / absent present., 93 'Mesosoma, Pretergites Ill to V1 with stridulatory granules' / absent present., 94 'Metasoma, Metasomal segments 1-1V, dorsoventrally compressed' / absent present., 95 'Metasomal segments 1-1V, carinae' / paired_ventrosubmedian_carinae single_ventromedian_carina., 96 'Metasomal segments 1-1V, carinae' / 'more strongly developed on 111-IV than 1-11' 'more strongly developed on 1-11 than 111-IV.', 97 'Metasomal segment 1, ventrosubmedian carinae with circular configuration' / absent present., 98 'Metasomal segment V, ventrolateral carinae' / continuous_from_proximal_to_distal_edges 'discontinuous, interrupted in distal region.', 99 'Metasomal segment V, ventromedian carina distal portion' / straight bifurcating breaking_up_into_numerous_granules, 100 'Metasomal segment V, transverse carina' / absent 'bothriurid type 1, discontinuous and merging proximally with ve' 'bothriurid type 11, continuous, not merging proximally with ven', 101 'Metasomal segment V, transverse carina' / absent 'diplocentrid type, partially developed' 'diplocentrid type, fully developed', 102 'Metasomal segment 11, ventral surface with numerous spiniform granules' / absent present., 103 'Metasomal segment V (male), dorsal surface with paired androvestigia (glands)' / absent present., 104 'Telson, Vesicle (male), dorsal surface with androvestigia (glands)' / absent single paired, 105 Aculeus / 'long, shallowly curved' 'very short, sharply curved.', 106 Subaculear_tubercle / absent distinct., 107 'Vesicle (male), laterally compressed' / absent present., 108 'Vesicle (female), laterally compressed' / absent present., 109 'Vesicle (male), elongated with pair of distal lobes' / absent present., 110 'Vesicle, anterodorsal lateral lobes (sensu San Martin and Gambardella, 1974)' / present absent., 111 'Vesicle, ventral surface' / with_two_or_more_longitudinal_granular_carinae_extending_toward 'with transverse row of granules at proximal edge, remainder smo' without_granules, 112 'Vesicle, ventral surface with semicircular carina' / absent present., 113 Venom_glands / complex simple., 114 Venom_pigment / opalescent reddish., 115 'Behaviour, Mesosomal percussion' / absent present., ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 ] [ . . . . . . . . . . . ] Bioculus_caboensis 1020000110100011101110100011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Bioculus_comondae 2020000110100011101110100011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Bothriurus_bonariensis 1000100020100011101110100130101100000000021011003-000001110000000001100100-0001112111100000000010001000100000000000 Bothriurus_coriaceus 1000100020100011101110100130101100000000021011003-000001110000000001100100-0001112111100000000010001000100000000000 Brachistosternus_alienus 1000000021100011101110100130101100000000021011003-0000011100000001010--200-00011121111000010000-0000001000000000000 Brachistosternus_ehrenbergii 1000000021100011101110100130101100000000022011003-0000011100000001010--200-00011121111000010000-0000001000000000000 Brachistosternus_ferrugineus 1000000021100011101110100130101100000000021011004-0000011100000001010--200-00011121111000010000-0000001000000000000 Cazierius_gundlachii 1020000010100011101110100011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000000010 Cazierius_scaber 1020000010100011101110100011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000000010 Centromachetes_obscurus 1000000021100011101110100130101120000000021011003-000001110000000001200100-0001112111100000000010000000000000000000 Centromachetes_pocockii 1000000021100011101110100130101120000000021011003-000001110000000001200100-0001112111100000000010000000000000000000 Centruroides_gracilis 000000000000000000000000000---0-00000010000000--1---0000000---0000000--0-0-00000000000-0000000000000000000000000000 Cercaphonius_queenslandae 1000000021100001101110100130101020000000021011003-0000011100000000011000-0-0001112111100000000010010000000000000000 Cercophonius_squama 1000000021100001101110100130101020000000021011003-0000011100000000011000-0-0001112111100000000010010000000000000000 Chaeritus_granosus 200000001000001000000000000-000-00000020011001000---0000000---0000000--0-0-00000010000-0000000000000000000000000100 Cheloctonus_crassimanus 1111001110000000101110000010110-0010000002101100210100101011101000002000-0-1110202110000000000000000000010110120100 Cheloctonus_jonesii 1111001110000000101110000010110-0010000002101100210100101011101000002000-0-1110202110000000000000000000010110120100 Chiromachetes_fergusoni 112100111000000111111000001011??10?1100002101100210100101011101000022000-0-111?2??????0???000000000000001011?120100 Chiromachetes_tirupati 112100111000000111111000001011??10??000002101100210100101011101000022000-0-111?2??????0???000000000000001011?120100 Chiromachus_ochropus 1121001110000001111110000010110-1011100002101100210100101011101000022300-0-1110202110000000000000000000010110120100 Didymocentrus_hasethi 1020000110100011101010010030100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Didymocentrus_lesueurn 1020000110100011101010010030100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Diplocentrus_gertschi 1020000110100011101010000021100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Diplocentrus_mexicanus 1020000010100011101010000021100-0000000012101100200300001011011020002000-0-0110202110001--0000010000200011000010010 Habibiella_gaillardi 1121001110000000111000000010000-1000000002102101211300001011011000002100-0-0110202110000000000100000000010101100100 Hadogenes_tityrus 1111001110000001111110101010110-10110000022021004-0120-01001101000002000-0-1110102110000010000000000010010110120100 Hadogenes_troglodytes 1111001110000001111110101010110-10101000022021004-0120-01001101000002000-0-1110102110000010000000000010010110100100 Hemiscorpius_lepturus 1121001110000000111000000010000-1000000002101101211300001011011000002100-0-0110202110000000000100000000010101100100 Hemiscorpius_maindroni 1121001110000000111000000010000-1000000002101101211300001011011000002100-0-0110202110000000000100000000010101100100 Heterometrus_spinifer 1021001010100011101111100011100-01100000121011002012000010110110100220011210110202110000000000000000000000000100000 Heterometrus_swammerdami 1021001010100011101111100011100-01100000121011002012000010110110100220011210110202110000000000000000000000000100000 Heteronebo_granti 1020000010100010101000000011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000100011000000010 Heteronebo_jamaicae 1020000010100010101000000011100-0000000012101100200300001011011020002000-0-0110202110001--0000010000100011000000010 Heteroscorpion_goodmani 2021001110100000111000000010000-20000000022021004-0000001001011000002000-0-0110202110000000000100000000000110100100 Heteroscorpion_opisthacanthoides 2021001110100000111000000010000-20000000022021004-0000001001011000002000-0-0110202110000000000100000000000110100100 Iomachus_laeviceps 1121001130000001111110000010110-101?000002101100210100101011101000002200-0-1110202110000100000000000000010110120100 Iomachus_politus 1121001130000001101110000010110-1020010002101100210100101001101000002200-0-1110202110000100000000000000010110120100 Liocheles_australasiae 1121001130000001111110000010110-1011000002101100210101101001101000022200-0-1110202110000100000000000010010110120100 Liocheles_waigiensis 1121001130000001111110000010110-1011000002101100210101101001101000022200-0-1110202110000100000000000000010110120100 Lisposoma_elegans 1000000011100000101110100130100-0000000002111100200000011100001000012000-0-00002021110001000000000-0000000000020000 Lisposoma_josehermana 1000000011100000101110100130100-0000000002111100200000011100001000012000-0-00002021110001000000000-0000000000020000 Nebo_hierichonticus 1021001010100001101110000010000-0100000002101110200300001011011010002000-0-0110202110001--0000000000000011000000010 Nebo_poggesii 1021001010100001101110000010000-0100000002101110200300001011011010002000-0-0110202110001--0000000000000011000000010 Oiclus_purvesii 2020000010100011101110100011100-0000000012101100200300001011011120002000-0-0110202110001--0000010000200011000000010 Opisthacanthus_elatus 1121001110000001111110000010110-1021010002101100210000101011101000002000-0-1110202110000100000000000000010110120100 Opisthacanthus_madagascariensis 1121001110000001111110000010110-1010100002101100210100101011101000002000-0-1110202110010000000000000000010110120100 Opisthacanthus_validus 1111001110000000101110000010110-1010{01}00002101100210100101011101000002000-0-1110202110000000000000000000010110100100 Opistophfhalmus_boehmi 1011001010101111101111000011100-010000001210210020120000101101101002201100-0110202110000000000010020000000000100000 Opistophthalmus_capensis 1011011010101111101111000011100-010000001210210020120000101101101002201200-0110202110000000000000000000000000100000 Opistophthalmus_holmi 1011011010101011101111000011100-01000000122021004-120000101101101002201200-01102021100000000000000-0000000000100000 Orobothriurus_alticola 1000100021100011101110100130101100000000021011003-000001110000000001200100-0001112111100000000010000000100000000000 Orobothriurus_crassimanus 1000100021100011101110100130101100000000021011003-000001110000000001200100-0001112111100000000010001000100000000000 Palaeocheloctonus_pauliani 1121001110000001111110000010110-0010100002101100210100101011101000002000-0-1110202110010000000000000000010110120100 Pandinus_cavimanus 1021001010100011101111100011100-01100000123022004-12000010110110100220011200110202110000000000000000000000000100000 Pandinus_dictator 1021001010100011101111100011100-01100000123021002012000010110110100220011200110202110000000000000000000000000100000 Pandinus_imperator 1021001010100011101111100011100-01100000123022004-12000010110110100220011200110202110000000000000000000000000100000 Phoniocercus_piclus 1000000021100001101110100130101000000000021011003-0000011100001000012200-0-00011121111000000000000-0000000000000000 Phoniocercus_sanmartini 1000000021100001101110100130101000000000021011003-0000011100001000012200-0-00011121111000000000000-0000000000000000 Scorpio_maurus_kruglovi 1011001010100011101111000011100-01000001121011002012000010110110100220010100110202110000000000010020000000000100001 Scorpio_maurus_mogadorensis 1011001010100011101111000011100-01000001121011002012000010110110100220010100110202110000000000010020000000000100001 Tarsoporosus_flavus 1020000010100011101000000011100-0000000012101100200300001011011020102000-0-0110202110001--0001010000200011000000010 Tehuankea_moyanoi 1000000021100001101110100130101100000000021011003-000001110000000001100??0-000?1121111000000000-0000000000000000000 Thestylus_glasioui 1000000021100001101110100130100-0000000002101100200000011100001000012000-0-00001121111000000000110-0000000000020000 Timogenes_dorbignyi 1000100020100011101110100130102-00000000021011004-000001110000000001100200-00011121111000000100-01-2001000000001000 Timogenes_mapuche 1000100020100011101110100130102-00000000022021004-000001110000000001100200-00011121111000000100-01-2001200000001000 Torsoporosus_kugleri 1020000010100011101000000011100-0000000012101100200300001011011020102000-0-0110202110001--0001010000200011000000010 Urodacus_novaehollandiae 2021001010100000101000000010000-20000000022021004-0010-0100101101002200100-0110202110000000100100010000000000100000 Urodacus_yaschenkoi 2021001010110000101000000010000-20000000022021004-0010-0100101101002200200-0110202110000000100100010000000000100000 Urophonius_granulatus 1000000021100001101110100130101020000000021011003-0000011100000000011000-0-0001112111100000000010010000100000000000 Urophonius_iheringii 1000000021100001101110100130101020000000021011003-0000011100000000011000-0-0001112111100000000010010000100000000000 Vachonia_martinezi 2000100020100011101110100130102-00000010023021004-000001110000100001100200-000?1121111000000000-01-2000000000000000 ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M2638] TITLE Table_4; LINK TAXA = Taxa2; DIMENSIONS NCHAR=47; FORMAT DATATYPE=Standard SYMBOLS= "0 1" MISSING=? GAP= -; CHARSTATELABELS 1 '(26). Carapace Median longitudinal furrow (a)' / 'broad, shallow (0)' 'suturiform (1).', 2 '(27). Median longitudinal furrow (b)' / 'suturiform (0)' 'broad, shallow-reversal (1).', 3 '(28). Sternum Shape (a)' / 'subpentagonal (0)' 'subtriangular (1).', 4 '(29). Sternum Shape (b)' / 'subpentagonal (0)' 'transverse (1).', 5 '(30). Sternum Shape (c)' / 'subpentagonal (0)' 'equilateral pentagonal (1).', 6 '(31). Chelicerae, Number of subdistal teeth' / '1 (0)' '2 (1).', 7 '(43). Pedipalp Ornamentation, Chela, male (a)' / 'no secondary sexual structures (0)' 'with secondary sexual structures (1).', 8 '(44). Chela, male (b)' / 'with hook-like structure (0)' 'with rimmed depression (1).', 9 '(45). Number of primary finger rows' / 'single (0)' 'multiple (1).', 10 '(49). Trichobothria, Femur, number of e t.' / 'one (0)' 'two or more (1).', 11 '(51). Patella, position of t d2' / 'dorsal (0)' 'internal (1).', 12 '(52). Patella, v t (a)' / 'absent (0)' 'three or more retrolaterals present (1).', 13 '(53). Patella, v t (b)' / 'all ventral (0)' 'one or more positioned externally (1).', 14 '(60). Patella, e t' / '7 (0)' '13 or more (1).', 15 '(69). Chela, number of i t.' / 'two (0)' 'one (1).', 16 '(70). Chela, position of r it' / 'basal (0)' 'distal (1).', 17 '(72). Chela, positions of i t.' / 'basal (0)' 'midfingerhemiscorpiines (1).', 18 '(75). Chela, number of V t.' / 'four (0)' 'five or more (1).', 19 '(79). Chela, distance between t V2 and V3' / 'normal (0)' 'widely separated (1).', 20 '(84). Chela, position of t-Dt (b)' / 'basal (0)' 'distalscorpionoids (1).', 21 '(85). Chela, position of t. DC-(c)' / 'distal (0)' 'midpalm (1).', 22 '(86). Chela, position of t- Est' / 'distal (0)' 'midpalm (1).', 23 '(87). Chela, position of t Et2' / 'external (0)' 'ventral (1).', 24 '(90). Legs, Retrolateral pedal spurs' / 'present (0)' 'absent-scorpionoids (1).', 25 '(92). Laterodistal lobes of telotarsi' / 'truncated (0)' 'rounded (1).', 26 '(93). Ventrosubmedian setae on telotarsi (a)' / 'setiform (0)' 'spiniform (1).', 27 '(94). Ventrosubmedian setae on telotarsi (b)' / 'spiniform (1)' 'secondarily setiform (1).', 28 '(95). Stridulatory organ' / 'mostly lacking (0)' 'on coxae of first leg and pedipalp (1).', 29 '(96). Length of first pair of maxillary lobes' / 'equal to second pair (0)' 'longer than second pair (1).', 30 '(101). Reproductive Anatomy, Type of embryonic development' / 'apoikogenic (0)' 'katoikogenic (1).', 31 '(103). Position of follicles on ovariuterus' / 'sessile (0)' 'stalked (1).', 32 '(104). Testis' / 'straight (0)' 'coiled (1).', 33 '(105). Genital opercula, female (a)' / 'separated (0)' 'loosely joined (1).', 34 '(107). Genital opercula, female (c)' / 'loosely joined (0)' 'fused (1).', 35 '(109). Genital opercula, male' / 'separated (0)' 'loosely joined (1).', 36 '(110). Hemispermatophore and Paraxial Organ, Hernispermatophore type (a)' / 'fusiform (0)' 'flagelliform (1).', 37 '(111). Hemispermatophore type (b)' / 'fusiform (0)' 'lamelliform (1).', 38 '(114). Truncal flexure (a)' / 'lacking (0)' 'mostly developed (1).', 39 '(117). Mating plug (a)' / 'gelatinous (0)' 'sclerotised (1).', 40 '(120). Internobasal reflection of sperm duct' / 'lacking (0)' 'well developed (1).', 41 '(121). Internal outer wall of sperm duct' / 'simple (0)' 'with semilunar shelf (1).', 42 '(122). Distal lamina' / 'smooth (0)' 'with prominent crest (1).', 43 '(123). Lamellar hook and median lobe' / 'separate (0)' 'fused (1).', 44 '(132). Metasoma, Ventral keels of segments I-IV (a)' / 'paired (0)' 'single-scorpionoids (1).', 45 '(134). Transverse ventral keel on segment V' / 'lacking (0)' 'well developed (1).', 46 '(135). Subaculear tubercle' / 'lacking (0)' 'well developed (1).', 47 '(136). Venom gland type' / 'complex (0)' 'simple (1).', ; MATRIX [ 10 20 30 40 ] [ . . . . ] Bioculus 11000000001101000001000101000110110011010010110 Bothriurus 00010010000101000100001001000001101011011100100 Brachistosternus 00010110000101000100001001000001101011011100000 Buthidae 00100000010000100000000000000000000100000000000 Cazierius 11000000001101000001000101000110110011010010110 Centromachetes 00010110100101000100001001000001101011011100000 Cercophonius 00010110100101000100001001000001101011011100000 Chaerilus 00000000010000000000000000000000000000000000000 Cheloctonus 10000000000101000011110101001110110011010000001 Chiromachetes 1000000010010100001111010100111011001?0???00001 Chiromachus 10000000100101000011110101101110110011010000001 Didymocentrus 11000000001101000001000101000110110011010010110 Diplocentrus 11000000001101000001000101000110110011010010110 Habibiella 10000000100101001011000101000110110011010001001 Hadogenes 100000001001010000?11?0101001110100011010000001 Hemiscorpius 10000000100101001011000101000110110011010001001 Heterometrus 10000000001101000001000111011110110011010000000 Heteronebo 11000000001101000001000101000110110011010010010 Heteroscorpion 100000000001010000?000010100?110110011010001001 Iomachus 10001000100101000011110101101110110011010000001 Liocheles 10001000100101000011110101101110110011010000001 Lisposoma 00010100000111000000001101000000100011011000000 Nebo 10000000000101010001000101000110110011010010010 Oiclus 11000000001101000001000101000110110011010010110 Opisthacanthus 10000000100101000011110101001110110011010000001 Opistophthalmus 10000000001101000001000111001110110011010000000 Orobothriurus 00010110000101000100001001000001101011011100000 Pandinus 10000000001101000001000111011110110011010000000 Phoniocercus 00010110000101000100001101000001101011011100000 Scorpio 10000000001101000001000111001110110011010000000 Tarsoporosus 11000000001101000001000101000110110011010010110 Tehuankea 0001011000010100010000100100000?101011011100000 Thestylus 0001010000010100000000110100000?101011011100000 Timogenes 00010011000101000100001001000001101011011100100 Urodacus 100000001001010000?0000111000110110011110001000 Urophonius 00010110100101000100001001000001101011011100000 Vachonia 0001001100010100010000110100000?101011011100100 ; END; BEGIN TREES; TITLE Tb5750; LINK TAXA = Taxa2; TRANSLATE 1 Centromachetes, 2 Bothriurus, 3 Chaerilus, 4 Buthidae, 5 Diplocentrus, 6 Didymocentrus, 7 Cazierius, 8 Bioculus, 9 Vachonia, 10 Brachistosternus, 11 Urophonius, 12 Timogenes, 13 Thestylus, 14 Tehuankea, 15 Phoniocercus, 16 Orobothriurus, 17 Lisposoma, 18 Cercophonius, 19 Urodacus, 20 Scorpio, 21 Pandinus, 22 Opistophthalmus, 23 Heterometrus, 24 Hemiscorpius, 25 Habibiella, 26 Opisthacanthus, 27 Liocheles, 28 Iomachus, 29 Hadogenes, 30 Chiromachus, 31 Chiromachetes, 32 Cheloctonus, 33 Heteroscorpion, 34 Nebo, 35 Tarsoporosus, 36 Oiclus, 37 Heteronebo; TREE Fig._1a = [&R] ((((((((35,36,7,8,6,5),37),34),((21,23),22,20)),19),(((((28,27),30),31,32,26,29),(24,25)),33)),((((10,((9,12),2),(11,18,1),14,16),15),13),17)),3,4); END; BEGIN TREES; TITLE Tb5751; LINK TAXA = Taxa1; TRANSLATE 1 Didymocentrus_lesueurn, 2 Didymocentrus_hasethi, 3 Chiromachus_ochropus, 4 Chiromachetes_fergusoni, 5 Chiromachetes_tirupati, 6 Cheloctonus_jonesii, 7 Cheloctonus_crassimanus, 8 Chaeritus_granosus, 9 Cercophonius_squama, 10 Cercaphonius_queenslandae, 11 Centruroides_gracilis, 12 Centromachetes_pocockii, 13 Centromachetes_obscurus, 14 Cazierius_scaber, 15 Cazierius_gundlachii, 16 Brachistosternus_ferrugineus, 17 Vachonia_martinezi, 18 Urophonius_iheringii, 19 Urophonius_granulatus, 20 Urodacus_yaschenkoi, 21 Urodacus_novaehollandiae, 22 Torsoporosus_kugleri, 23 Timogenes_mapuche, 24 Timogenes_dorbignyi, 25 Thestylus_glasioui, 26 Tehuankea_moyanoi, 27 Tarsoporosus_flavus, 28 Scorpio_maurus_mogadorensis, 29 Scorpio_maurus_kruglovi, 30 Phoniocercus_sanmartini, 31 Phoniocercus_piclus, 32 Pandinus_imperator, 33 Pandinus_dictator, 34 Pandinus_cavimanus, 35 Palaeocheloctonus_pauliani, 36 Orobothriurus_crassimanus, 37 Orobothriurus_alticola, 38 Opistophthalmus_holmi, 39 Opistophthalmus_capensis, 40 Opistophfhalmus_boehmi, 41 Opisthacanthus_validus, 42 Opisthacanthus_madagascariensis, 43 Opisthacanthus_elatus, 44 Oiclus_purvesii, 45 Nebo_poggesii, 46 Nebo_hierichonticus, 47 Iomachus_politus, 48 Lisposoma_josehermana, 49 Lisposoma_elegans, 50 Liocheles_waigiensis, 51 Liocheles_australasiae, 52 Iomachus_laeviceps, 53 Heteroscorpion_opisthacanthoides, 54 Heteroscorpion_goodmani, 55 Heteronebo_jamaicae, 56 Heterometrus_swammerdami, 57 Heterometrus_spinifer, 58 Heteronebo_granti, 59 Hemiscorpius_maindroni, 60 Hemiscorpius_lepturus, 61 Hadogenes_troglodytes, 62 Hadogenes_tityrus, 63 Bioculus_caboensis, 64 Habibiella_gaillardi, 65 Diplocentrus_mexicanus, 66 Diplocentrus_gertschi, 67 Brachistosternus_ehrenbergii, 68 Brachistosternus_alienus, 69 Bothriurus_coriaceus, 70 Bothriurus_bonariensis, 71 Bioculus_comondae; TREE Fig._2 = [&R] (((((((((((2,1)Didymocentrus,66),65),((58,55)Heteronebo,(27,22)Tarsoporosus)),((63,71)Bioculus,15,14,44))Diplocentrinae,(46,45)Nebinae)Diplocentridae,((((32,34),33)Pandinus,(57,56)Heterometrus),(((39,38),40)Opistophthalmus,(29,28)Scorpio_maurus))Scorpionidae),((((((((((51,50)Liocheles,47),52),43),((4,3),5)),(42,35)),(62,61)Hadogenes),(7,6)Cheloctonus),41)Ischnuridae,(64,60,59)Hemiscorpiidae)),((21,20)Urodacidae,(54,53)Heteroscorpionidae)),((((((((((((24,23)Bothriurinae,17),70,69),36),37),(13,12)Centromachetes),(67,68,16)Brachistosterninae),26),((18,19)Urophonius,10,9)),(31,30)Phoniocercus),25),(49,48)Lisposoma)Bothriuridae),8,11)Scorpiones; END; BEGIN TREES; TITLE Tb5752; LINK TAXA = Taxa1; TRANSLATE 1 Didymocentrus_lesueurn, 2 Didymocentrus_hasethi, 3 Chiromachus_ochropus, 4 Chiromachetes_fergusoni, 5 Chiromachetes_tirupati, 6 Cheloctonus_jonesii, 7 Cheloctonus_crassimanus, 8 Chaeritus_granosus, 9 Cercophonius_squama, 10 Cercaphonius_queenslandae, 11 Centruroides_gracilis, 12 Centromachetes_pocockii, 13 Centromachetes_obscurus, 14 Cazierius_scaber, 15 Cazierius_gundlachii, 16 Brachistosternus_ferrugineus, 17 Vachonia_martinezi, 18 Urophonius_iheringii, 19 Urophonius_granulatus, 20 Urodacus_yaschenkoi, 21 Urodacus_novaehollandiae, 22 Torsoporosus_kugleri, 23 Timogenes_mapuche, 24 Timogenes_dorbignyi, 25 Thestylus_glasioui, 26 Tehuankea_moyanoi, 27 Tarsoporosus_flavus, 28 Scorpio_maurus_mogadorensis, 29 Scorpio_maurus_kruglovi, 30 Phoniocercus_sanmartini, 31 Phoniocercus_piclus, 32 Pandinus_imperator, 33 Pandinus_dictator, 34 Pandinus_cavimanus, 35 Palaeocheloctonus_pauliani, 36 Orobothriurus_crassimanus, 37 Orobothriurus_alticola, 38 Opistophthalmus_holmi, 39 Opistophthalmus_capensis, 40 Opistophfhalmus_boehmi, 41 Opisthacanthus_validus, 42 Opisthacanthus_madagascariensis, 43 Opisthacanthus_elatus, 44 Oiclus_purvesii, 45 Nebo_poggesii, 46 Nebo_hierichonticus, 47 Iomachus_politus, 48 Lisposoma_josehermana, 49 Lisposoma_elegans, 50 Liocheles_waigiensis, 51 Liocheles_australasiae, 52 Iomachus_laeviceps, 53 Heteroscorpion_opisthacanthoides, 54 Heteroscorpion_goodmani, 55 Heteronebo_jamaicae, 56 Heterometrus_swammerdami, 57 Heterometrus_spinifer, 58 Heteronebo_granti, 59 Hemiscorpius_maindroni, 60 Hemiscorpius_lepturus, 61 Hadogenes_troglodytes, 62 Hadogenes_tityrus, 63 Bioculus_caboensis, 64 Habibiella_gaillardi, 65 Diplocentrus_mexicanus, 66 Diplocentrus_gertschi, 67 Brachistosternus_ehrenbergii, 68 Brachistosternus_alienus, 69 Bothriurus_coriaceus, 70 Bothriurus_bonariensis, 71 Bioculus_comondae; TREE Fig._6 = [&R] ((((((((((((51,50),47),52),43),((4,3),5)),(((7,6),41),(62,61)),(42,35)),(64,60,59)),54,53),(21,20)),((((63,71),15,14,44),(((2,1),66),65),((58,55),(27,22))),(46,45)),((((32,34),33),(57,56)),(((39,38),40),(29,28)))),(((((((((((24,23),17),70,69),36),37),(67,68,16),(13,12)),26),((18,19),10,9)),(31,30)),25),(49,48))),8,11); END;