#NEXUS [!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file. Generated on April 24, 2024; 13:51 GMT TreeBASE (cc) 1994-2008 Study reference: Wilson G.D., & Edgecombe G. 2003. The Triassic isopod Protamphisopus wianamattensis (Chilton) and comparison with extant taxa (Crustacea, phreatoicidea). Journal of Paleontology, 77(3): 454-470. TreeBASE Study URI: http://purl.org/phylo/treebase/phylows/study/TB2:S920] BEGIN TAXA; TITLE Taxa1; DIMENSIONS NTAX=61; TAXLABELS Amphisopus_annectans Amphisopus_lintoni Colubotelson_fontinalis Colubotelson_joyneri Colubotelson_searli Colubotelson_sp._1 Colubotelson_sp._2 Crenisopus_acinifer Crenoicus_buntiae Crenoicus_harrisoni Crenoicus_n.sp. Eophreatoicus_sp._4 Eophreatoicus_sp._6 Eremisopus_beei Gariwerdeus_beehivensis Gariwerdeus_ingletonensis Gariwerdeus_turretensis Hyperoedesipus_plumosus Hypsimetopus_intrusor Kalliapseudes_obtusifrons Mesacanthotelson_setosus Mesacanthotelson_tasmaniae Mesamphisopus_abbreviatus Mesamphisopus_capensis Mesamphisopus_n.sp. Metaphreatoicus_affinis Metaphreatoicus_australis Metaphreatoicus_lacustris Metaphreatoicus_magistri Metaphreatoicus_sp. Naiopegia_xiphagrostis Neophreatoicus_assimilis Nichollsia_kashiense Notamphisopus_dunedinensis Notamphisopus_flavius Onchotelson_brevicaudatus Onchotelson_spatulatus Paramphisopus_palustris Paraphreatoicus_relictus Peludo_paraliotus Phreatoicoides_gracilis Phreatoicoides_sp._A Phreatoicoides_sp._B Phreatoicoides_sp._Zeehan Phreatoicopsis_raffae Phreatoicopsis_terricola Phreatoicus_orarii Phreatoicus_typicus Phreatomerus_latipes Pilbarophreatoicus_n.sp. Pilbarophreatoicus_platyarthricus Platypyga_subpetrae Protamphisopus_wianamattensis Qld._nsp.1 Qld._nsp.2 Spelaeogriphus_lepidops Stenasellus_virei Synamphisopus_ambiguus Synamphisopus_doegi Tainisopus_fontinalis Uramphisopus_pearsoni ; END; BEGIN CHARACTERS; [! TreeBASE Matrix URI: http://purl.org/phylo/treebase/phylows/matrix/TB2:M410] TITLE Phreatoicidea; LINK TAXA = Taxa1; DIMENSIONS NCHAR=165; FORMAT DATATYPE=Standard SYMBOLS= "1 2 3 4 5 6" MISSING=? GAP= -; CHARSTATELABELS 1 Cervical_groove / present absent, 2 'Mandibular (genal or cheek) groove' / 'smoothly indented, or with acute indentation' absent, 3 Mandibular_notch / present absent, 4 Clypeal_notch / present absent, 5 Antennal_notch / present absent, 6 'Pereonites 2-7 in dorsal view (relative lengths)' / longer_than_wide wider_than_long anteriorly_longer_than_wide_decreasing_posteriorly_to_wider_than_long, 7 'Coxal articulation of pereonites 2-4' / free nearly_fused_or_fused, 8 'Coxal articulation of pereonites 5-7' / free nearly_fused_or_fused, 9 Sternal_processes_on_sternites / present absent, 10 'Typhlosole (form in cross-section)' / 'well developed, ventral invagination forming double spiral in cross section' 'intermediate, ventral invagination forming Y shape in cross section' 'minimal, ventral invagination forming laminar projection or inverted u shape in cross section' 'absent, gut round in cross section', 11 'Pleonites in lateral view (depth of ventrolateral plates relative to pereonites)' / 'much deeper than pereonites, with large pleurae, basal region of pleopods not visible' 'having depth approximating depth of pereonites, with small pleurae, basal region of pleopods visible', 12 Pleonites_with_pleonite_1_pleura / 'distinctly shallower than pleurae of pleonites 2-5' 'near depth of pleurae of pleonites 2-5', 13 Pleonite_5_ventral_margin_shape / 'subparallel to dorsal margin along length, not sinuate posteriorly' sinuate, 14 Pleotelson_form / 'shallow (lateral fields sloping)' 'vaulted (lateral fields vertical)', 15 Pleotelson_dorsal_surface_shape_in_lateral_view / evenly_curving strongly_inflected_ventrally, 16 Pleotelson_dorsal_surface_setae / lacking_simple_setae sparsely_covered_with_fine_setae covered_with_abundant_curled_simple_setae covered_with_abundant_elongate_setae densely_covered_with_cuticular_hairs_and_setae with_abundant_robust_setae, 17 Pleotelson_dorsal_surface_median_ridge / absent present, 18 Pleotelson_dorsal_surface_lateral_ridges / continuous_with_posterior_apex absent, 19 Pleotelson_lateral_length_versus_depth / less_than_depth_or_subequal_to_depth greater_than_depth, 20 Pleotelson_ventral_surface_anterior_to_uropods / strongly_concave 'flattened, only slightly concave', 21 Pleotelson_ventral_margin_anterior_to_uropods_robust_setae / with_robust_setae with_distally_denticulate_setae_in_one_row with_distally_denticulate_robust_setae_in_two_rows with_fine_setae_only, 22 Pleotelson_ventral_margin_relative_lengths_of_robust_setae_anterior_to_uropods / posterior_seta_longer_than_anterior_adjacent_setae posterior_seta_length_subequal_to_anterior_adjacent_setae posterior_seta_shorter_than_anterior_adjacent_setae, 23 Pleotelson_postanal_ventral_surface / 'present (apical margin and anus separated by cuticular surface)' 'absent (anus adjacent to apical margin)', 24 Pleotelson_postanal_ventral_surface_ornamentation / 'unelaborated, or with fine setae on ridge' 'with robust setae (not on ridge)' with_robust_setae_on_ridge with_groove_forming_cleft_in_posterior_apex, 25 Pleotelson_lateral_uropodal_ridge / terminating_at_pleotelson_margin_above_uropods curving_strongly_and_extending_posteriorly_from_uropods absent, 26 Pleotelson_lateral_uropodal_ridge_setae / with_row_of_robust_setae with_1_robust_seta with_fine_simple_setae lacking_setae, 27 'Pleotelson posterolateral margin, differentiation from apex' / uninterrupted with_distinct_inflection_differentiating_apex, 28 Pleotelson_posterolateral_margin_shape / forming_rounded_process_with_dorsally_projecting_setae crenate forming_vertical_plate forming_rounded_lobe unelaborated, 29 'Pleotelson posterolateral margin (shape if lobe-like)' / curving_dorsally depressed_ventrally_and_posteriorly_angular, 30 'Pleotelson posterolateral margin robust setae (if plates or rounded lobes presen' / without_robust_setae with_one_position_with_robust_setae with_two_positions_with_robust_setae_with_intermediate_setae with_two_positions_with_robust_setae_without_intermediate_setae with_multiple_postions_with_robust_setae_with_intermediate_setae, 31 'Pleotelson posterolateral margin dorsal robust setae (if plates or lobes present' / with_one_robust_seta 'with multiple robust setae, ventrolateral seta' 'with multiple robust setae, ventrolateral setae subequal to dorsomedial setae', 32 'Pleotelson posterolateral margin submarginal seta near inflection point (if plat' / present absent, 33 'Pleotelson posterolateral margin dorsolateral surface robust setae (if lobe-like' / absent adjacent_to_margin set_anteriorly_from_margin, 34 'Pleotelson posterior apex shape in dorsal view (if postanal ventral surface pres' / projecting indented transverse, 35 'Pleotelson posterior apex (flattened or not)' / 'free (not flattened)' strongly_reflexed_and_flattened_against_dorsal_surface, 36 'Pleotelson posterior apex projections (if postanal ventral surface absent)' / with_small_rounded_tubercle without_projections, 37 'Pleotelson posterior apex length in lateral view(if projecting)' / 'very long (> 0.295 pleotelson length)' 'long (0.120-0.295 pleotelson length)' 'short (< 0.119 pleotelson length)', 38 'Pleotelson posterior apex shape in dorsal view (if protruding freely)' / elongate broadly_rounded quadrate, 39 Pleotelson_posterior_apex / visible_in_lateral_view obscured_by_posterolateral_margin_in_lateral_view, 40 Pleotelson_posterior_apex_robust_setae / absent with_1_pair_of_robust_setae with_2_pairs_of_robust_setae with_3_pairs_of_robust_setae with_4_pairs_of_robust_setae, 41 'Antennula articles (in male)' / more_than_6_articles 6_or_fewer_articles, 42 Antennula_article_3_rudimentary_second_flagellum / present absent, 43 'Antennula article 4 length (relative to article 3)' / shorter 'subequal (ratio 0.95-1.05)' longer, 44 'Antennula terminal article shape (taxa with more 6 articles in male)' / tubular globular 'distally oblique, with 2 or more groups of aesthetascs' conical 'tiny, vestigial', 45 'Antennula terminal article length relative to penultimate article (taxa more tha' / shorter subequal longer, 46 Antennula_penultimate_article_length / longer_than_any_other_article 'length approximately sub-equal to, or less than length of other articles', 47 'Antennula articles 5-6 shape (taxa with 6 or fewer articles in male)' / inflated_and_bulbous width_subequal_to_article_4 article_5_only_inflated_and_bulbous, 48 'Antennula article 5 length relative to article 6 (taxa with 6 or fewer articles' / subequal_or_less greater article_6_not_expressed, 49 'Antennula penultimate and ante-penultimate articles shape in cross section' / flattened 'oval, or circular', 50 'Antenna flagellum proximal articles dense cover of cuticular hairs (in male)' / present absent, 51 'Antenna flagellum proximal articles dense cover of cuticular hairs (in female)' / present absent, 52 Antenna_propodal_article_3_scale / well_developed rudimentary_but_articulated absent, 53 Antenna_article_5_length_relative_to_article_4 / shorter_or_subequal longer, 54 Antenna_article_6_length_relative_to_articles_4_and_5_combined / shorter_or_subequal longer, 55 Mandible_palp_third_article_shape / 'crescent-like' relatively_linear, 56 'Mandible palp third article setae on medial-distal margins' / with_fewer_than_5_setae with_more_than_5_setae, 57 Mandible_palp_3rd_article_coarsely_spinulate_setae / absent coarsely_spinulate_setae_with_other_setae coarsely_spinulate_setae_present_only, 58 Mandible_palp_third_article_medial_surface / naked covered_in_dense_cuticular_hairs covered_in_cuticular_combs, 59 Mandible_incisor_processes_width_relative_to_thickness / broad thin, 60 Mandible_right_lacinia_mobilis_shape / 'large, distinctly separated' 'indistinct, spine-like' absent, 61 Mandible_right_lacinia_mobilis_dentate_plates / two_dentate_plates single_dentate_plate, 62 Mandible_spine_row_bifurcate_spines / present absent, 63 Mandible_spine_row_general_shape_between_incisor_and_molar / in_smooth_low_arc on_projecting on_linear_pedunculate_projection on_round_pedunculate_projection on_elongate_penicil, 64 Mandible_spine_rows_shape_of_spine_ridge_in_ventral_view / distal_margin_in_line_with_proximal_margin distal_margin_protruding_in_relative_to_proximal_margin 'forming strongly convex arc, protruding medially', 65 Mandible_spine_rows_shape_of_spine_ridge_in_medial_view / basal_insertions_in_line_between_incisor_and_molar_processes basal_insertions_crossing_dorsally_and_then_abruptly_angling_posteriorly, 66 Mandible_right_spine_row_first_spine_relative_to_remaining_spines / separated not_separated, 67 Maxillula_medial_lobe_width / less_than_or_subequal_to_lateral_lobe greater_than_lateral_lobe, 68 Maxillula_medial_lobe_pappose_setae / up_to_three_setae four_setae five_setae six_setae_or_more, 69 Maxillula_medial_lobe_accessory_setae / one_or_two more_than_two absent, 70 Maxillula_lateral_lobe_distal_margin_denticulate_robust_setae / with_one_or_more absent, 71 Maxillula_lateral_lobe_distal_margin_smooth_robust_setae / up_to_nineteen twenty_or_more, 72 Maxillula_lateral_lobe_ventral_face_setae_position / widely_spaced closely_spaced, 73 '(Maxillula) lateral lobe ventral face plumose seta' / present absent, 74 Maxilla_outer_lateral_lobe_distal_margin_setal_row / approximately_linear_and_transverse_to_lobe_axis curving_and_extending_proximally_along_medial_margin 'with two angles - transverse to lateral margin and oblique on medial margin', 75 'Pereopod I dactylus ventrodistal margin cuticular fringe (in male)' / with_multiple_rows_of_minute_sharp_spines with_row_of_sharp_spines 'with row of thin scale-like spines' 'with cuticular fringe (minute cuticular hairs)' cuticular_fringe_absent, 76 'Pereopod I dactylus ventrodistal margin cuticular fringe (in female)' / with_multiple_rows_of_minute_sharp_spines with_row_of_sharp_spines 'with row of thin scale-like spines' cuticular_fringe_absent, 77 Pereopod_I_dactylus_accessory_claws / several one_only absent, 78 Pereopod_I_dactylus_distal_accessory_spines / several one_only absent, 79 'Pereopod I propodus dorsal margin setae (in male)' / setae_present_in_several_groups_between_proximal_and_distal_margin setae_confined_to_single_group_at_distal_margin, 80 'Pereopod I propodus dorsal margin proximal region (in male)' / protruding_to_distodorsal_margin_of_ischium protruding_to_distodorsal_margin_of_carpus not_protruding, 81 'Pereopod I propodus dorsal margin proximal region (in female )' / protruding_to_distodorsal_margin_of_ischium protruding_to_distodorsal_margin_of_carpus not_protruding, 82 'Pereopod I propodal palm shape (in male)' / concave convex_to_straight, 83 'Pereopod I propodal palm proximal simple spine (in male)' / present absent, 84 'Pereopod I propodal palm median simple spine (in male)' / present absent, 85 'Pereopod I propodal palm distal simple spines (in male)' / one more_than_one absent, 86 'Pereopod I propodal palm medial rugose cuticular pad (in male)' / present absent, 87 'Pereopod I propodal palm stout denticulate setae (in male)' / serrate bifid absent, 88 'Pereopod I propodal palm stout robust setae (in male)' / conical basally_inflated absent, 89 'Pereopod I propodal palm setal ridge (in male)' / absent present, 90 'Pereopod I propodal palm shape (in female)' / concave convex straight, 91 'Pereopod I propodal palm midpoint simple spine (in female)' / present absent, 92 'Pereopod I propodal palm stout denticulate setae (in female)' / serrate bifid absent, 93 'Pereopod I propodal palm stout robust simple setae (in female)' / conical basally_inflated absent, 94 'Pereopod I merus distodorsal margin in anterodorsal view (in male)' / 'shelf-like and U-shaped' 'spine-like and pointed', 95 'Pereopod I merus distodorsal margin projection in anterodorsal view (in female)' / 'shelf-like and U-shaped' 'spine-like and pointed', 96 'Pereopod I merus distodorsal margin setal form (in male)' / with_more_than_2_elongate_simple_setae 'with 1-2 robust simple setae' 'with 1-2 elongate simple setae', 97 'Pereopods II-III propodus articular plate' / present absent, 98 'Pereopod II basis dorsal margin proximal group of setae (in male)' / robust_setae elongate_fine_setae without_distinct_group_of_large_setae, 99 'Pereopod II basis dorsal margin proximal group of setae (in male)' / in_1_row 'in 2 rows (rarely more)', 100 'Pereopod II basis dorsal margin projection (in male)' / present absent, 101 'Pereopod II basis dorsal margin proximal group of setae (in female)' / robust_setae elongate fine setae proximally without_distinct_group_of_large_setae, 102 'Pereopod II basis dorsal margin proximal group of setae number of rows (in femal' / in_1_row 'in 2 rows (rarely more)', 103 'Pereopod II basis dorsal margin projection (in female)' / present absent, 104 'Pereopod IV prehensility (sexual dimorphism, adult male form)' / prehensile 'similar to female, not prehensile', 105 'Pereopod IV subchelate major hinges (in male if dimorphic)' / on_dactylus_and_propodus on_propodus_and_carpus, 106 'Pereopod IV dactylus length (in male)' / shorter_than_or_subequal_to_propodal_palm longer_than_propodal_palm, 107 Pereopod_IV_propodus_articular_plate / present absent, 108 'Pereopods V-VII propodus articular plate' / present absent, 109 'Pereopods V-VII basis dorsal ridge shape' / rounded_or_angular_on_all forming_distinct_plate_at_least_on_pereopod_VII, 110 'Pereopods V-VII basis dorsal margin' / with_robust_setae with_elongate_fine_setae without_large_setae, 111 'Pereopods V-VII basis lateral face central longitudinal ridge' / present absent, 112 'Pereopods V-VII basis lateral face ventral longitudinal ridge' / present absent, 113 'Pereopods V-VII basis lateral face ventral longitudinal ridge' / with_fine_setae setae_absent, 114 Pereopod_VII_ischium_dorsal_ridge_flange_width / greater_than_or_subequal_to_shaft_width less_than_shaft_width flange_absent, 115 Pereopod_VII_basis_dorsal_ridge_distal_margin_shape / indented rounded, 116 Penes_form / straight curved_posteriorly, 117 Penes_extent_medially / near_or_reaching_midline past_midline_and_onto_first_pleonite, 118 Penes_shaft_cuticle / smooth with_cuticular_hairs denticulate, 119 Penes_setae / on_shaft on_tip absent, 120 Penes_distal_tip_shape / rounded pointed flattened truncate, 121 'Pleopod exopods II-V proximal article distolateral lobes length relative to dist' / shorter_than_distal_article subequal_or_longer_than_distal_article, 122 'Pleopod exopods I-V lateral proximal lobes' / 'on I-V' 'on II-V' absent_on_II absent, 123 'Pleopod exopods I-V medial proximal lobes' / 'on I-V' 'on II-V' 'on III-V' absent, 124 'Pleopod endopods marginal setation (in male)' / 'endopods I-V without setae' endopod_I_only_with_setae 'endopods I-V with setae', 125 Pleopod_protopods_medial_margins / 'I-V with epipods or small projections' 'II-V with epipods or small projections' 'III-V with epipods' 'I-V without epipods', 126 'Pleopod I-V protopods medial margin coupling hooks (in male)' / 'present (on at least some pleopods)' absent, 127 'Pleopod protopods medial margin setae (in male)' / with_plumose_setae with_minutely_serrate_or_simple_setae setae_absent, 128 'Pleopod protopods medial margin coupling hooks (in female)' / present absent, 129 'Pleopod protopods medial margin pleopod I coupling hook numbers (in female)' / one_only with_multiple_coupling_hooks, 130 Pleopod_protopod_I_lateral_epipod / 'lobe-like' linear absent, 131 Pleopod_protopod_II_lateral_epipod / 'lobe-like' linear absent, 132 'Pleopod protopods lateral epipods III-V' / 'lobe-like' absent, 133 Male_pleopod_I_exopod_lateral_margin_shape / rounded broadly_angular concave convoluted, 134 Male_pleopod_I_exopod_dorsal_surface / with_setae lacking_setae, 135 Male_pleopod_I_exopod_ventral_surface_shape / flat curving_anteriorly, 136 Male_pleopod_I_protopod_shape / longer_than_wide width_subequal_length significantly_wider_than_long, 137 'Male pleopod II endopod appendix masculina (stylet) proximal shape ventrallly' / broadly_concave 'rod-like, convex' 'rod-like, indented', 138 Male_pleopod_II_endopod_stylet_channel / tube_along_distal_length not_forming_tube, 139 Male_pleopod_II_endopod_stylet_tip_shape / truncateor_broadly_rounded acutely_rounded conical 'sharply pointed and spine-like', 140 Male_pleopod_II_endopod_stylet_tip_denticles / smooth_or_with_minute_rounded_denticles with_large_subterminal_denticles, 141 Male_pleopod_II_endopod_stylet_marginal_setation / multiple_setae 1_seta setae_absent, 142 Male_pleopod_II_endopod_stylet_length_relative_to_exopod / near_to_distal_margin not_extending_beyond_half_length, 143 Uropod_protopod_extension_posteriorly_in_lateral_view / beyond_or_subequal_to_pleotelson_apex not_reaching_pleotelson_apex, 144 Uropod_protopod_dorsomedial_ridge_shape / 'plate-like' 'spur-like' rounded_projection, 145 Uropod_protopod_dorsomedial_ridge_shape / parallel_to_ventral_margin laterally_concave_at_midpoint, 146 Uropod_protopod_dorsomedial_ridge_margin_setal_form / robust_spinose_distally robust_simple fine_simple absent, 147 Uropod_protopod_distoventral_margin_robust_setal_number / absent one two three four_or_more, 148 Uropod_protopod_distoventral_margin_robust_spinose_setae / absent present, 149 Uropod_rami_distal_tips_shape / pointed rounded, 150 'Uropod rami cross-sectional shape' / round flattened_dorsally_and_ventrally flattened_on_dorsal_surface_only 'exopod round, endopod flattened', 151 Uropod_endopod_length_relative_to_protopod / shorter subequal longer, 152 Uropod_endopod_length_relative_to_exopod / subequal_or_longer distinctly_shorter, 153 Uropod_endopod_shape / straight_or_curving_dorsally curving_medially, 154 Uropod_endopod_dorsal_margin_robust_setae_placement / all_along_length midlength proximally absent, 155 Uropod_endopod_spine_on_dorsal_margin / present absent, 156 Uropod_endopod_proximoventral_margin_shape_in_medial_view / sinuate convex, 157 'Uropod exopod robust setae on dorsal margin (in male)' / one_or_more absent, 158 'Uropod exopod robust setae on dorsal margin (in female)' / one_or_more absent, 159 Pereopod_II_basis_proximal_dorsal_ridge / absent present, 160 Pereopodal_exopods / present absent, 161 Short_carapace / present absent, 162 Heart_position / thoracic 'pleonal-thoracic', 163 Uropodal_articles / either_multiarticulate uniarticulate, 164 Telson / free fused_to_pleonite_6, 165 Antennular_second_flagellum / multiarticulate tiny_or_absent, ; MATRIX [ 10 20 30 40 50 60 70 80 90 100 110 120 130 140 150 160 170 180 190 ] [ . . . . . . . . . . . . . . . . . . . ] Amphisopus_annectans 211112212411-212211112141123-521-31---12121312--2223212211111222221311121122231331---23311-3311113--2111221123112312223412212121-331422211311111124223311221111222222 Amphisopus_lintoni 211112212411-212211111141123-521-31---12121322--2223212211111121?21411121322331331---23311-3311113--2121221123112312213312212121-331422211313111124223311221111222222 Colubotelson_fontinalis 111112212311-223121123111324122-211-2213121212--2223112221121131121211121153231332---21113-1311112123--111111?21231211{12}112212222-33112123-111113124211311222111222222 Colubotelson_joyneri 112112222211-222121121111324122-211-221312121{12}--122321222112?13112121212113323?332---21113-1311112122121111122212312111112212222-33111123-111113124214111422221222222 Colubotelson_searli 112212222411-22?12112111132412?--11-221312121{12}--1??321??2?12?13112121?1???3332?332---21113-111111212212111112222-312111112212222-33112123-1111?3124214111222111222222 Colubotelson_sp._1 111112212411-222121123111324122-211-2213121212--2223212221121131121211121133231332---21112-1311112122121111112212312211112212222-33111123-111113123211311222111222222 Colubotelson_sp._2 111112212411-221121122111324122-211-2213121212--12231122111211311212111211??23?332---21113-1311112122121111121212312211112212222-33111123-111113122211311422111222222 Crenisopus_acinifer 122222212411-212121112112425-----11-3311221---222223122111111122121111121?5423122212323311-3321323--3--1112211212311113112212222-331111212211113114223211222111222222 Crenoicus_buntiae 121112112311-22312111{23}111324121-311-3213121212--122311??1?121131111211122122211232---23121-311111212212111111222-312213112212222-33121122-412113122211211422111222222 Crenoicus_harrisoni 121112112311-22?12111{12}1?132412?--11-3?13121212--122311??1?12?131111211121?32221212---23123-3111112122121-1111222-312213112212222-33121122-4121?3122211211222111222222 Crenoicus_n.sp. 121112112311-22312111{23}111324121-311-3213121212--12231122111211311212111--154231222---23123-311111212212111111222--?2213112212222-33121122-412113122211211{24}22121222222 Eophreatoicus_sp._4 111112{12}12411-22222111{12}132423-412-11-222312?312--2223??2211111121221411121352231222---23112-1111311112211111123112122223{13}11112222-311121212111111124123311222112222222 Eophreatoicus_sp._6 111112{12}12411-222221111132423-412-11-2223121312--222321221311112221141112135423?222---23112-111131221221111112311212221{12}111112222-311121212111111124123311222112222222 Eremisopus_beei 111112212311-212211121112423-421-11-1212121312--2113212211111122221211122353{12}31232---23112-3111313--3--1111123112222113112232121-311121212111111123213311222111222222 Gariwerdeus_beehivensis 111212212311-22412112211132421--111-331{23}221---11222311221112113112121112113323133{12}---23111-3111113--3--1111112211312213112212222-33111123-211113123211311422221222222 Gariwerdeus_ingletonensis 111212212311-22412112211132421--111-3311221---232223112211121131121211121333231331---21311-13111122122111111122113122131122122{12}2-33112123-111113225211311422221222222 Gariwerdeus_turretensis 111212212311-22412112311132421--111-3311221---112223{12}12211121131121211121154232232---23211-1111112122121111112211312213112212222-33112123-111113232211311422221222222 Hyperoedesipus_plumosus {12}2112222242212141222422-3-12-------2----121511--22232122111111211212111-2354231{12}31---23313-3311123--212111221322-3111134{12}14121211332121112211112111-23111422221222222 Hypsimetopus_intrusor 12112?????22121{23}122?1???1415------------1?1???-----?2122??1{23}?1{23}?111{34}?11????????3?1---?3?1??????????????2--??1????31?????21?2?????3311?11??{234}1?2{12}3?2??1?1112??1?1222222 Kalliapseudes_obtusifrons -----211--???11?122???113-?5-----1?-----1111?2-----12?????13-25?121??21????????-3?---?331?-33---1??????2--??1????3-----?-4-342?2-3321212----?--3--1-21?11?22??1111121 Mesacanthotelson_setosus 111112212311-222121121111125-----11-1113121212--222311222112?131?21211121153232332---21312-1311112122121111112211312233412212222-33112123-111113123211{23}11222111222222 Mesacanthotelson_tasmaniae 111112212?11-22{26}122123111125-----11-1113121212--2??3{12}1??2?1{23}?131121{23}111???5223?332---21113-131111111112111111322-3122{13}3112212222-33112123-1111?3123211311222111222222 Mesamphisopus_abbreviatus 111112112?11-222121111112415-----11---12{12}2?512122??3????1?111123??12211????233?331---2??11-{12}3?111???21211?112?21231?????12232121-3311?12121?11?11?4123311{23}22111222222 Mesamphisopus_capensis 111112112411-222121111112415-----11---12{12}11512122223{12}2??1?111123221211111332231332---22211-211111212212111112321231211{12}112232121-331111212111111124123311222111222222 Mesamphisopus_n.sp. 121112112411-222121111112415-----11---1?121512--2??311??1?111123221211?---3?23?332---22211-2?1111??????1111123112312111112232121-331121?12111111124122311222111222222 Metaphreatoicus_affinis 111112212311-222121122111324122-211-22131?1212--2223212221121131121211121353231332---21112-1311113--3--1111112212312223112212222-33112123-111113124211311222111222222 Metaphreatoicus_australis 111112212311-223121122111324122-211-221{34}121212--2223112211121131121211121153231232---2{13}113-1311112112111111122211312122112212222-33112123-111113123211311222111222222 Metaphreatoicus_lacustris 111112212311-222121123111324122-211-2213121212--222311221112113?121?11121153231232---21113-1311112112111111122211312123112212222-33112123-111113123211311222111222222 Metaphreatoicus_magistri 111112212311-223121123111324122-211-2213121212--2223111221121131121311121133231332---21312-13111121221211111{12}2211312122112212222-33111123-111113124211311{34}22111222222 Metaphreatoicus_sp. 111112212311-223121123111324122-211-2213121212--12?321??2?1{23}1131121211121?53231332---21113-131111212211111112?211312123412212222-33112123-1111?3123211311222111222222 Naiopegia_xiphagrostis 111112212311-222121112111324121-111-2213221---222113112211121131121211121122231331---23111-3111113--3--1111112211312213112212222-33112123-111113224211311422221222222 Neophreatoicus_assimilis 1?211311??11-21?1211??1?112??????11---1?121212-----322??1?221133??121??????????2?22132312????1??1??????1111?{12}????3122??112?14222-3311?113-11{12}21312311?111?22??1222222 Nichollsia_kashiense 112121221422?2121222122-3-12-------2----121122---22{13}2122111112232214111---2233?331---23111-3111323--3--2--221322-311113422412222-332121212211212141-{12}4122422221222222 Notamphisopus_dunedinensis 121112212?11-2{12}212112{12}111{123}24122-211-221{34}121{12}1{12}--2??3{12}1????1{23}?13?1?12111-{12}??????232---13123-311111??????11111122113121???122{12}222??33112113-2111?3124211211422221222222 Notamphisopus_flavius 121112212311-223121121111324131-211-221212121{12}--1223212211121131121{23}111-1133231232---13123-3111113--3--111111221131211{12}412222222-33112113-111113124211{23}11{24}22221222222 Onchotelson_brevicaudatus 111112212311-222121133111325-----11-1112121122--2113{12}12231121131121211121153232332---21313-1311113--3--111111222-312223412212222-33111113-111111122211311222221222222 Onchotelson_spatulatus 111112212311-223121133111325-----11-1112121112--21??2122311211311?1211121153232332---21313-1311113--3--111111222-312213412212222-33111113-111111122211311222221222222 Paramphisopus_palustris 212222222411-211121111121123-521-11-221{345}121312--2213212211111123221411121352231332---23113-3311111122112--1123212322123313212111-331111211313111122123211222111222222 Paraphreatoicus_relictus 111112212311-222121122111324122-211-2213121212--1223112221121131121{23}11121133231332---21312-131111211211111111221231222{12}1122{23}2222-33111123-1111?3123211311222111222222 Peludo_paraliotus 111112211411-225121122111423-1-2-12---11121312--2113212212111122221211121333132331---23111-3111113--3--2--112312-2122233111222{12}2-231321312111111141-11311422221222222 Phreatoicoides_gracilis 12112121242222121222122-3-15-------1----121512--2223212211111121121211122?54331331{12}2{13}23311-3311123-----2--221222-312113421412222-332121112311212111-14{12}22412221222222 Phreatoicoides_sp._A 12112221142222221222232-3-15-------1----121512--122321????111121121{12}11122353231331---23311-3311123--2122--221222-312213{34}22412222-332121112311213111-14322312221222222 Phreatoicoides_sp._B 12112321142212111222122-3-15-------1----121512--1223212211111121121311121353231331{12}2323311-3311223--2122--221222-312213{34}21412222-33212111231?213121-11311222111222222 Phreatoicoides_sp._Zeehan 122122222422121?1222112-1315------------121412-----321??1?12?122?212111???5413?23112123311-3122323--2122--221222-3121114{12}1412222-3321211123131?2111-11112212221222222 Phreatoicopsis_raffae 211112211111-2121211132-2415-------2----111312--2223212112112141-214222122543312222132311313122323--3--2--2211112312213311111222-22112123-221121123113311222111222222 Phreatoicopsis_terricola 211112211111-2121211132-2415-------2----111312--2223212112112141-21422212254331222213231131312232??????2--2213112312{12}13311111222-22112123-221121123113311222111222222 Phreatoicus_orarii 122213212?11-22212211?1?1425-----11-3213121212-----321??1?22?13112241???????23?232---23121-31111???????111??1?22-312213112312222-33111123-11{12}1?3122211111222111222222 Phreatoicus_typicus 122213212211-222122111113-25-----11-3213121212--122321??1?22?132122411122?22212232---23121-3111113--3--1-1111222-311213112212222-33112113-111113122114111222111222222 Phreatomerus_latipes 211112222411-212121112142{13}23-521-11-221{45}121312--222321221111112222141112131{13}232232---23313-3311113--2122--1123112222213212312111-331411211313111125113211222111222222 Pilbarophreatoicus_n.sp. 21222121142212121222122-3-12-------2----121511---223212211111122?2141?1---332322211212331123311311121122--111322-3111134224142{12}2-33211111231{13}213121-23111422221222222 Pilbarophreatoicus_platyarthricus 1?122?21?42212121222122-3-12-------2----121511---22321?????11122?21{12}?11???5423?2?11212331????1?313--???2--111322-31?113?22414222-3321211????3213121-2?211422221222222 Platypyga_subpetrae 211112212411-222121113111421-----22-----121322--2223212211111122221211122?32231232---23211-1311{13}121121111{12}11231123122131122121312331121212211113122123211222111222222 Protamphisopus_wianamattensis 1???1?11??11-21?1211??1?1?15-----31---111?111?--2???212??????????????????????????1???????1?????????????2--??2?1??32???????????????????????????231???1?311?22??2?2?22? Qld._nsp.1 111122212411-213121111112415-----21-----123512--22231112111121232211311-135123133111223311-3311123--3--112{12}123212322211{34}21213222-332121212211213125123311122111222222 Qld._nsp.2 121122212411-213121111112421-----21-----123512--22231222?1?1??????11111-135{23}23123211223321-33?1123--3--112222222-322211{23}21211222-331121212{12}11113125223311222111222222 Spelaeogriphus_lepidops -----211--???11?122???113-?5-----1?-----1111?2-----12?????11121?121??21????????33?---?331?-33-1-???????2--??1????3-1113?-4-341?1-3321212----?--3--1-22?21?22??1111111 Stenasellus_virei ?????211?4???11?12????113-?5-----1?-----1211?2--2??121????13-211121???????2????332---?131?-13---1??????2--111????3---13?14-142?2-332--12----?--3--1-21311?22??1222222 Synamphisopus_ambiguus 211112212111-222221112112421-----21-----111322--2223212213112141-212122123332322322{12}321312-1111321121121112121112312213312211222-3311212123211111{12}4223311222111222222 Synamphisopus_doegi 211112212111-222221112112421-----21-----111312--2223212213112141-21112212333231232---23212-1311323--21211121211123122123122112{12}2-331121212321111123223311222111222222 Tainisopus_fontinalis ?2211211?4???11?12????113-?5-----1?-----1111?2--2??221????111211121???????4????332---?131?-13---1??????2--111????3---13?14-211?1-11112131221?1?3--1-22111?22??1222222 Uramphisopus_pearsoni 111112212411-222121121111325-----11-2114121212--222321222?121131121211121154232232---21312-131111222222111111222-312223412212222-33111123-111111123211311222111222222 ; END; BEGIN TREES; TITLE Tb6317; LINK TAXA = Taxa1; TRANSLATE 1 Protamphisopus_wianamattensis, 2 Platypyga_subpetrae, 3 Pilbarophreatoicus_platyarthricus, 4 Pilbarophreatoicus_n.sp., 5 Phreatomerus_latipes, 6 Phreatoicus_typicus, 7 Phreatoicus_orarii, 8 Phreatoicopsis_raffae, 9 Phreatoicopsis_terricola, 10 Phreatoicoides_gracilis, 11 Phreatoicoides_sp._Zeehan, 12 Phreatoicoides_sp._B, 13 Phreatoicoides_sp._A, 14 Peludo_paraliotus, 15 Paraphreatoicus_relictus, 16 Paramphisopus_palustris, 17 Onchotelson_spatulatus, 18 Onchotelson_brevicaudatus, 19 Notamphisopus_flavius, 20 Notamphisopus_dunedinensis, 21 Nichollsia_kashiense, 22 Neophreatoicus_assimilis, 23 Metaphreatoicus_sp., 24 Metaphreatoicus_magistri, 25 Metaphreatoicus_lacustris, 26 Metaphreatoicus_australis, 27 Metaphreatoicus_affinis, 28 Mesamphisopus_n.sp., 29 Mesamphisopus_capensis, 30 Mesamphisopus_abbreviatus, 31 Mesacanthotelson_tasmaniae, 32 Mesacanthotelson_setosus, 33 Hypsimetopus_intrusor, 34 Hyperoedesipus_plumosus, 35 Eremisopus_beei, 36 Eophreatoicus_sp._6, 37 Eophreatoicus_sp._4, 38 Crenoicus_n.sp., 39 Crenoicus_harrisoni, 40 Crenoicus_buntiae, 41 Crenisopus_acinifer, 42 Colubotelson_sp._2, 43 Colubotelson_sp._1, 44 Colubotelson_searli, 45 Colubotelson_joyneri, 46 Colubotelson_fontinalis, 47 Amphisopus_lintoni, 48 Amphisopus_annectans, 49 Stenasellus_virei, 50 Tainisopus_fontinalis, 51 Kalliapseudes_obtusifrons, 52 Spelaeogriphus_lepidops, 53 Qld._nsp.2, 54 Qld._nsp.1, 55 Gariwerdeus_ingletonensis, 56 Gariwerdeus_beehivensis, 57 Gariwerdeus_turretensis, 58 Naiopegia_xiphagrostis, 59 Uramphisopus_pearsoni, 60 Synamphisopus_doegi, 61 Synamphisopus_ambiguus; TREE Fig._14 = [&R] (52,(51,((50,49),((((((((48,47),(16,5)),((((37,36),35),2),((30,29),28))),(((((46,(32,31,(18,17),59),((26,25),23),24),((45,44),42),43,15),27),((((40,39,38),(22,(7,6))),(20,19)),(58,((57,55),56)))),14)),(((9,8),1),(61,60))),(54,53)),41),(34,((((33,12),13),10),11),(21,(4,3))))))); END;