#NEXUS

[!This data set was downloaded from TreeBASE, a relational database of phylogenetic knowledge. TreeBASE has been supported by the NSF, Harvard University, Yale University, SDSC and UC Davis. Please do not remove this acknowledgment from the Nexus file.


Generated on May 26, 2024; 8:34 GMT

TreeBASE (cc) 1994-2008

Study reference:
Portik D.M. 2019. Sexual Dichromatism Drives Diversification Within a Major Radiation
of African Amphibians. Systematic Biology, .

TreeBASE Study URI:  http://purl.org/phylo/treebase/phylows/study/TB2:S23981]

BEGIN TAXA;
        TITLE  Untitled_Block_of_Taxa;
        DIMENSIONS NTAX=283;
        TAXLABELS
            Acanthixalus_sonjae
            Acanthixalus_spinosus
            Afrixalus_brachycnemis
            Afrixalus_delicatus
            Afrixalus_dorsalis_1
            Afrixalus_dorsalis_2
            Afrixalus_dorsalis_3
            Afrixalus_dorsimaculatus_1
            Afrixalus_dorsimaculatus_2
            Afrixalus_enseticola
            Afrixalus_equatorialis
            Afrixalus_fornasini
            Afrixalus_fulvovittatus_1
            Afrixalus_fulvovittatus_2
            Afrixalus_knysae
            Afrixalus_lacteus
            Afrixalus_lacustris
            Afrixalus_laevis
            Afrixalus_morerei
            Afrixalus_nigeriensis
            Afrixalus_osorioi
            Afrixalus_paradorsalis_1
            Afrixalus_paradorsalis_2
            Afrixalus_paradorsalis_3
            Afrixalus_quadrivittatus_1
            Afrixalus_quadrivittatus_2
            Afrixalus_spinifrons
            Afrixalus_stuhlmanni
            Afrixalus_sylvaticus
            Afrixalus_vibekensis
            Afrixalus_vittiger_1
            Afrixalus_vittiger_2
            Afrixalus_weidholzi
            Afrixalus_wittei
            Arthroleptis_adelphus
            Arthroleptis_affinis
            Arthroleptis_bioko
            Arthroleptis_brevipes
            Arthroleptis_fichika
            Arthroleptis_francei
            Arthroleptis_nikeae
            Arthroleptis_palava
            Arthroleptis_perreti
            Arthroleptis_poecilonotus
            Arthroleptis_reichei
            Arthroleptis_schubotzi
            Arthroleptis_stenodactylus
            Arthroleptis_sylvaticus
            Arthroleptis_taeniatus
            Arthroleptis_tanneri
            Arthroleptis_variabilis
            Arthroleptis_xenodactyloides
            Astylosternus_batesi
            Astylosternus_diadematus
            Astylosternus_laticephalus
            Astylosternus_occidentalis
            Astylosternus_schioetzi
            Balebreviceps_hillmani
            Breviceps_adspersus
            Breviceps_branchi
            Breviceps_fuscus
            Breviceps_gibbosus
            Breviceps_macrops
            Breviceps_montanus
            Breviceps_mossambicus
            Breviceps_namaquensis
            Callulina_dawida
            Callulina_hanseni
            Callulina_kanga
            Callulina_kisiwamsitu
            Callulina_kreffti
            Callulina_laphami
            Callulina_meteora
            Callulina_shengena
            Callulina_sp_Rubeho
            Callulina_sp_lowland
            Callulina_stanleyi
            Cardioglossa_annulata
            Cardioglossa_elegans
            Cardioglossa_escalerae
            Cardioglossa_gracilis
            Cardioglossa_leucomystax
            Cardioglossa_manengouba
            Cardioglossa_melanogaster
            Cardioglossa_nigromaculata
            Cardioglossa_occidentalis
            Cardioglossa_oreas
            Cardioglossa_pulchra
            Cardioglossa_trifasciata
            Chlorolius_koehleri
            Cryptothylax_greshoffii
            Hemisus_marmoratus
            Hemisus_sudanensis
            Heterixalus_alboguttatus
            Heterixalus_andrakata
            Heterixalus_betsileo
            Heterixalus_boettgeri
            Heterixalus_carbonei
            Heterixalus_luteostriatus
            Heterixalus_madagascariensis
            Heterixalus_punctatus
            Heterixalus_rutenbergi
            Heterixalus_tricolor
            Heterixalus_variabilis
            Hyperolius_acuticeps
            Hyperolius_ademetzi
            Hyperolius_adspersus
            Hyperolius_argus
            Hyperolius_balfouri_1
            Hyperolius_balfouri_2
            Hyperolius_baumanni
            Hyperolius_benguellensis
            Hyperolius_bobirensis
            Hyperolius_bocagei
            Hyperolius_bolifambae
            Hyperolius_burgessi
            Hyperolius_camerunensis
            Hyperolius_castaneus
            Hyperolius_chlorosteus
            Hyperolius_cinereus
            Hyperolius_cinnamomeoventris_1
            Hyperolius_cinnamomeoventris_2
            Hyperolius_cinnamomeoventris_3
            Hyperolius_concolor_1
            Hyperolius_concolor_2
            Hyperolius_concolor_3
            Hyperolius_cystocandicans
            Hyperolius_dartevellei
            Hyperolius_dintelmanni
            Hyperolius_discodactylus
            Hyperolius_endjami_1
            Hyperolius_endjami_2
            Hyperolius_friedemanni
            Hyperolius_frontalis
            Hyperolius_fusciventris_1
            Hyperolius_fusciventris_2
            Hyperolius_guttulatus_1
            Hyperolius_guttulatus_2
            Hyperolius_howelli
            Hyperolius_igbettensis
            Hyperolius_inyangae
            Hyperolius_jackie
            Hyperolius_jacobseni
            Hyperolius_kivuensis_1
            Hyperolius_kivuensis_2
            Hyperolius_kivuensis_3
            Hyperolius_kuligae
            Hyperolius_lamottei
            Hyperolius_langi
            Hyperolius_lateralis
            Hyperolius_lupiroensis
            Hyperolius_mariae
            Hyperolius_marmoratus_marmoratus
            Hyperolius_marmoratus_swynnertoni
            Hyperolius_marmoratus_taeniatus
            Hyperolius_minitissimus
            Hyperolius_mitchelli
            Hyperolius_molleri
            Hyperolius_montanus
            Hyperolius_mosaicus
            Hyperolius_nasutus
            Hyperolius_nitidulus_1
            Hyperolius_nitidulus_2
            Hyperolius_occidentalis
            Hyperolius_ocellatus_1
            Hyperolius_ocellatus_2
            Hyperolius_parallelus_alborufus
            Hyperolius_parallelus_marginatus
            Hyperolius_parallelus_parallelus
            Hyperolius_parallelus_pyrrhodictyon
            Hyperolius_pardalis
            Hyperolius_parkeri
            Hyperolius_phantasticus
            Hyperolius_picturatus
            Hyperolius_pictus
            Hyperolius_platyceps
            Hyperolius_poweri
            Hyperolius_pseudoargus
            Hyperolius_pusillus
            Hyperolius_quinquevittatus
            Hyperolius_raymondii
            Hyperolius_reesi
            Hyperolius_riggenbachi
            Hyperolius_rubrovermiculatus
            Hyperolius_rwandae
            Hyperolius_semidiscus
            Hyperolius_sp_nov_1
            Hyperolius_sp_nov_2
            Hyperolius_spatzi
            Hyperolius_spinigularis
            Hyperolius_steindachneri
            Hyperolius_substriatus
            Hyperolius_sylvaticus
            Hyperolius_tanneri
            Hyperolius_thomensis
            Hyperolius_torrentis
            Hyperolius_tuberculatus
            Hyperolius_tuberilinguis
            Hyperolius_veithi
            Hyperolius_viridiflavus_destefanii
            Hyperolius_viridiflavus_glandicolor
            Hyperolius_viridiflavus_ngorongoriensis
            Hyperolius_viridiflavus_ommatostictus
            Hyperolius_viridiflavus_pantherinus
            Hyperolius_viridiflavus_pitmani
            Hyperolius_viridiflavus_sp
            Hyperolius_viridiflavus_variablilis
            Hyperolius_viridis
            Hyperolius_zonatus
            Kassina_arboricola
            Kassina_cassinoides
            Kassina_cochranae
            Kassina_decorata
            Kassina_fusca
            Kassina_kuvangensis
            Kassina_lamottei
            Kassina_maculosa
            Kassina_schioetzi
            Kassina_senegalensis
            Leptodactylodon_axillaris
            Leptodactylodon_bicolor
            Leptodactylodon_boulengeri
            Leptodactylodon_bueanus
            Leptodactylodon_erythrogaster
            Leptodactylodon_mertensi
            Leptodactylodon_ornatus
            Leptodactylodon_ovatus
            Leptodactylodon_perreti
            Leptodactylodon_polyacanthus
            Leptodactylodon_ventrimarmoratus
            Leptopelis_argenteus
            Leptopelis_aubryi
            Leptopelis_aubryioides
            Leptopelis_bocagii
            Leptopelis_boulengeri
            Leptopelis_brevirostris
            Leptopelis_calcaratus
            Leptopelis_fiziensis
            Leptopelis_flavomaculatus
            Leptopelis_gramineus
            Leptopelis_karissimbensis
            Leptopelis_kivuensis
            Leptopelis_macrotis
            Leptopelis_millsoni
            Leptopelis_modestus
            Leptopelis_natalensis
            Leptopelis_nordequatorialis
            Leptopelis_notatus
            Leptopelis_ocellatus
            Leptopelis_palmatus
            Leptopelis_parkeri
            Leptopelis_rufus
            Leptopelis_spiritusnoctis
            Leptopelis_susanae
            Leptopelis_vannutellii
            Leptopelis_vermiculatus
            Leptopelis_viridis
            Leptopelis_yaldeni
            Morerella_cyanophthalma
            Nyctibates_corrugatus
            Opisthothylax_immaculatus
            Paracassina_kounhiensis
            Paracassina_obscura
            Phlyctimantis_boulengeri
            Phlyctimantis_keithae
            Phlyctimantis_leonardi
            Phlyctimantis_maculata
            Phlyctimantis_verrucosus
            Phrynomantis_microps
            Probreviceps_durirostris
            Probreviceps_loveridgei
            Probreviceps_macrodactylus
            Probreviceps_rungwensis
            Probreviceps_sp_Kigogo
            Probreviceps_sp_Nguru
            Probreviceps_sp_Rubeho
            Probreviceps_sp_Udzungwa
            Probreviceps_uluguruensis
            Scotobleps_gabonicus
            Semnodactylus_wealii
            Spelaeophryne_methneri
            Tachycnemis_seychellensis
            Trichobatrachus_robustus
            ;
END;

BEGIN TREES;
      TITLE African_Amphibians_Calibs2_30M_con;
      LINK TAXA = Untitled_Block_of_Taxa;
      TREE TREE1 = [&R] (Phrynomantis_microps:76.61564484708043,(((Hemisus_sudanensis:2.6904126706775457,Hemisus_marmoratus:2.6904126706775457):49.177906641743746,(Balebreviceps_hillmani:41.853907437036526,(((Breviceps_mossambicus:9.909645656658938,Breviceps_adspersus:9.909645656658938):13.830123333671459,(Breviceps_montanus:17.508932518511315,(Breviceps_fuscus:15.395398756248369,(Breviceps_gibbosus:14.619909315876306,(Breviceps_macrops:10.011184237191586,(Breviceps_namaquensis:6.288685291850015,Breviceps_branchi:6.288685291850015):3.72249894534157):4.608725078884724):0.7754894403720627):2.1135337622629464):6.230836471819096):14.106327746298135,((Spelaeophryne_methneri:23.700721818337307,((Probreviceps_loveridgei:8.830638817239901,Probreviceps_uluguruensis:8.830638817239901):8.016869100306465,((Probreviceps_durirostris:5.1021371417295285,Probreviceps_sp_Nguru:5.1021371417295285):6.10733409768703,(Probreviceps_sp_Rubeho:9.830692935336657,(Probreviceps_sp_Kigogo:4.4952099078046075,(Probreviceps_sp_Udzungwa:2.467478026131886,(Probreviceps_rungwensis:1.0552684542012827,Probreviceps_macrodactylus:1.0552684542012827):1.412209571930589):2.027731881672736):5.335483027132042):1.378778305079905):5.638036677129804):6.853213900790941):5.549852909750605,((Callulina_shengena:10.971707636825613,Callulina_laphami:10.971707636825613):8.324756658161093,(Callulina_dawida:14.95623432216233,(Callulina_kanga:12.711682409699193,(Callulina_sp_lowland:12.186515883921224,(Callulina_sp_Rubeho:10.783468055115009,((Callulina_hanseni:5.344138773326236,Callulina_meteora:5.344138773326236):4.905288693918095,(Callulina_stanleyi:5.8030698446259805,(Callulina_kisiwamsitu:1.5046655375251419,Callulina_kreffti:1.5046655375251419):4.2984043070708395):4.446357622618336):0.5340405878706918):1.4030478290062192):0.5251665257779692):2.2445519124631375):4.340229972824375):9.954110432101203):8.59552200854062):4.007810696407994):10.014411875384766):20.27927011903475,(((Nyctibates_corrugatus:35.53191428944093,(Scotobleps_gabonicus:30.762574178430754,((Astylosternus_laticephalus:6.7036243056939355,Astylosternus_occidentalis:6.7036243056939355):16.00863609757473,(Trichobatrachus_robustus:16.096406852797386,(Astylosternus_diadematus:7.144937814037036,(Astylosternus_schioetzi:3.285952470039504,Astylosternus_batesi:3.285952470039504):3.8589853440975332):8.951469037760361):6.615853551471268):8.050313774162085):4.76934010901018):9.857439904414633,(((Leptodactylodon_mertensi:12.512937459313406,(Leptodactylodon_erythrogaster:8.915773735498462,(Leptodactylodon_perreti:3.710505889611781,Leptodactylodon_axillaris:3.710505889611781):5.205267845886695):3.59716372381493):11.57598758485259,((Leptodactylodon_polyacanthus:4.088525849142513,Leptodactylodon_bicolor:4.088525849142513):14.829611543419418,(Leptodactylodon_bueanus:13.371880814075908,(Leptodactylodon_ornatus:12.125354939329597,(Leptodactylodon_boulengeri:9.443425811156203,(Leptodactylodon_ventrimarmoratus:7.137033539743271,Leptodactylodon_ovatus:7.137033539743271):2.3063922710129248):2.6819291291734118):1.2465258747462968):5.546256577486034):5.170787650604062):18.97772128301022,((Leptopelis_parkeri:35.30674161100836,((Leptopelis_macrotis:7.845988286007824,Leptopelis_millsoni:7.845988286007824):17.92494995943577,(Leptopelis_rufus:21.349802184346572,((Leptopelis_argenteus:16.19204951456576,(Leptopelis_yaldeni:8.165976584177997,(Leptopelis_vannutellii:6.954350244337888,(Leptopelis_susanae:4.481300229739276,Leptopelis_gramineus:4.481300229739276):2.473050014498625):1.2116263398401088):8.026072929887754):3.150581007746176,((Leptopelis_kivuensis:15.64683796498511,(Leptopelis_ocellatus:12.84654552061859,(Leptopelis_spiritusnoctis:8.714440961555553,(Leptopelis_viridis:6.5126901690851255,Leptopelis_aubryi:6.5126901690851255):2.2017507925204356):4.132104558963036):2.800292444466521):1.5332041185003717,(Leptopelis_natalensis:16.146251317951986,((Leptopelis_palmatus:13.572180491547144,(Leptopelis_calcaratus:11.699231230841292,(Leptopelis_brevirostris:8.17180910368387,Leptopelis_notatus:8.17180910368387):3.5274221271174184):1.8729492608058536):1.8903452353889563,((Leptopelis_modestus:11.50372408364538,(Leptopelis_flavomaculatus:8.115556013000074,(Leptopelis_nordequatorialis:3.112318389336025,Leptopelis_bocagii:3.112318389336025):5.0032376236540586):3.388168070645307):2.583483295386408,((Leptopelis_vermiculatus:8.594066321437381,Leptopelis_aubryioides:8.594066321437381):3.280463470905147,(Leptopelis_boulengeri:9.759437113005745,(Leptopelis_fiziensis:4.715747106745141,Leptopelis_karissimbensis:4.715747106745141):5.043690006270609):2.1150926792367812):2.212677586789262):1.3753183480043134):0.6837255909158841):1.0337907656334977):2.1625884380264537):2.00717166303464):4.421136061097009):9.535803365564767):5.129312897898579,(((Cardioglossa_melanogaster:21.21115673842384,Cardioglossa_gracilis:21.21115673842384):3.8132659594685094,((Cardioglossa_elegans:16.42506853087972,(Cardioglossa_leucomystax:14.694416625188587,Cardioglossa_occidentalis:14.694416625188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.976142802683341,Afrixalus_dorsalis_1:6.976142802683341):2.221428750884357,((Afrixalus_quadrivittatus_1:5.239088732958066,Afrixalus_quadrivittatus_2:5.239088732958066):2.932418846747069,(Afrixalus_dorsalis_2:6.1095312767452725,(Afrixalus_fulvovittatus_2:4.541902618172557,Afrixalus_fulvovittatus_1:4.541902618172557):1.5676286586227235):2.061976302959863):1.026063973762561):1.0305103520978207):2.7234150035968483):0.6227046749490697):2.331069440408733):1.4144799955531795):3.125707201010904):4.088490821442321):7.503155647675932,((Morerella_cyanophthalma:17.431105654630827,Cryptothylax_greshoffii:17.431105654630827):13.396224976705398,(Hyperolius_semidiscus:28.21056288280893,((Hyperolius_lupiroensis:2.415713032301042,Hyperolius_parkeri:2.415713032301042):22.92328281317574,(((Hyperolius_viridis:12.055425767285108,Hyperolius_pusillus:12.055425767285108):3.098646690304612,((Hyperolius_benguellensis:8.399621618419957,Hyperolius_inyangae:8.399621618419957):3.7078279272927546,(Hyperolius_nasutus:9.69902700153014,(Hyperolius_dartevellei:9.300393248841942,(Hyperolius_rwandae:8.54789334352968,(Hyperolius_friedemanni:7.589160476770246,((Hyperolius_howelli:1.2444822290196385,Hyperolius_acuticeps:1.2444822290196385):6.028857544902365,((Hyperolius_poweri:5.215503109777643,Hyperolius_igbettensis:5.215503109777643):1.4033055403796482,(Hyperolius_jacobseni:2.7871016720365986,Hyperolius_adspersus:2.7871016720365986):3.8317069781206925):0.6545311237647127):0.31582070284824226):0.9587328667294344):0.7524999053122627):0.39863375268819823):2.408422544082569):3.0466229119769963):9.225427359781762,(((Hyperolius_pseudoargus:15.079451124186079,((Hyperolius_fusciventris_2:4.457692690617293,Hyperolius_fusciventris_1:4.457692690617293):9.77799706156378,(Hyperolius_bolifambae:9.948094030297696,((Hyperolius_ocellatus_1:4.264377682729531,Hyperolius_ocellatus_2:4.264377682729531):3.923676213919201,(Hyperolius_sp_nov_1:7.393557457148475,(Hyperolius_camerunensis:4.369282524301809,Hyperolius_riggenbachi:4.369282524301809):3.024274932846666):0.7944964395002572):1.7600401336889675):4.287595721883378):0.8437613720050052):2.09063013108765,((Hyperolius_guttulatus_2:4.577416628497275,Hyperolius_guttulatus_1:4.577416628497275):10.123636674285322,((Hyperolius_phantasticus:5.630769412639765,(Hyperolius_pardalis:3.591433681316687,Hyperolius_steindachneri:3.591433681316687):2.039335731323078):7.579301439664576,(Hyperolius_argus:12.514012009656447,(Hyperolius_bocagei:11.54020495036221,(Hyperolius_lamottei:9.577509237593304,(((Hyperolius_tuberculatus:4.67556065754016,Hyperolius_dintelmanni:4.67556065754016):1.8033668326060024,(Hyperolius_parallelus_parallelus:5.789057141294421,(Hyperolius_parallelus_marginatus:4.178259661594652,(Hyperolius_parallelus_alborufus:3.145162490837734,Hyperolius_parallelus_pyrrhodictyon:3.145162490837734):1.033097170760925):1.6107974796997553):0.6898703488557629):1.657498526748455,((Hyperolius_marmoratus_taeniatus:5.907952011083879,((Hyperolius_marmoratus_marmoratus:2.1957133353150766,Hyperolius_marmoratus_swynnertoni:2.1957133353150766):3.3254869473003623,(Hyperolius_mariae:2.956942308576288,Hyperolius_reesi:2.956942308576288):2.564257974039151):0.3867517284664359):1.8306105536781843,((Hyperolius_nitidulus_1:4.456782549399193,(Hyperolius_spatzi:3.2230221378057706,Hyperolius_nitidulus_2:3.2230221378057706):1.2337604115994196):2.740636563198052,((Hyperolius_viridiflavus_pitmani:5.355602865302103,(Hyperolius_viridiflavus_sp:4.729093813152417,(Hyperolius_viridiflavus_destefanii:2.7611245282180477,Hyperolius_viridiflavus_variablilis:2.7611245282180477):1.9679692849403665):0.6265090521426799):0.8144214098450391,(Hyperolius_viridiflavus_ngorongoriensis:5.215945667690093,(Hyperolius_viridiflavus_ommatostictus:3.539746755957509,(Hyperolius_viridiflavus_pantherinus:2.5070460264244616,Hyperolius_viridiflavus_glandicolor:2.5070460264244616):1.0327007295330475):1.6761989117345877):0.9540786074517484):1.0273948374551054):0.5411434521577974):0.397863452137571):1.4410832207036748):1.962695712818899):0.9738070591942503):0.6960588426478935):1.4909824503782545):2.469027952591148):5.477288863064771,(Chlorolius_koehleri:19.38454033103838,(Hyperolius_tuberilinguis:18.130012924223934,(((Hyperolius_mosaicus:4.01479487722068,(Hyperolius_endjami_2:2.090544805665374,Hyperolius_endjami_1:2.090544805665374):1.9242500715952957):8.451501783870242,(Hyperolius_cystocandicans:9.808835653563989,(Hyperolius_jackie:8.57748912896433,((Hyperolius_discodactylus:3.512908012465928,Hyperolius_lateralis:3.512908012465928):2.576490743730659,(Hyperolius_castaneus:4.571590905276821,Hyperolius_frontalis:4.571590905276821):1.5178078509097617):2.4880903727677577):1.2313465246296431):2.6574610075269334):4.564086492990128,(((Hyperolius_platyceps:10.16384757235491,(Hyperolius_cinereus:6.704799673715158,Hyperolius_raymondii:6.704799673715158):3.459047898649743):4.58516900113225,(Hyperolius_langi:9.793590038701453,(Hyperolius_kuligae:5.7401822032466185,Hyperolius_ademetzi:5.74018220324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[!  TreeBASE tree URI: http://purl.org/phylo/treebase/phylows/tree/TB2:Tr115835]


END;